Palm Nut Vulture
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The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in the family
Accipitridae The Accipitridae is one of the three families within the order Accipitriformes, and is a family of small to large birds with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects to medium-s ...
(which also includes many other diurnal
raptor Raptor or RAPTOR may refer to: Animals The word "raptor" refers to several groups of bird-like dinosaurs which primarily capture and subdue/kill prey with their talons. * Raptor (bird) or bird of prey, a bird that primarily hunts and feeds on v ...
s such as
kite A kite is a tethered heavier than air flight, heavier-than-air or lighter-than-air craft with wing surfaces that react against the air to create Lift (force), lift and Drag (physics), drag forces. A kite consists of wings, tethers and anchors. ...
s,
buzzard Buzzard is the common name of several species of birds of prey. ''Buteo'' species * Archer's buzzard (''Buteo archeri'') * Augur buzzard (''Buteo augur'') * Broad-winged hawk (''Buteo platypterus'') * Common buzzard (''Buteo buteo'') * Eastern ...
s and harriers, vultures, and eagles). It is the only member of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Gypohierax''. This bird is an
Old World vulture Old World vultures are vultures that are found in the Old World, i.e. the continents of Europe, Asia and Africa, and which belong to the family Accipitridae, which also includes eagles, buzzards, kites, and hawks. Old World vultures are not clos ...
(only distantly related to the
New World vulture The New World vulture or condor family, Cathartidae, contains seven extant species in five genera. It includes five extant vultures and two extant condors found in warm and temperate areas of the Americas. The "New World" vultures were widespread ...
s, which are in a separate family, the
Cathartidae The New World vulture or condor family, Cathartidae, contains seven extant species in five genera. It includes five extant vultures and two extant condors found in warm and temperate areas of the Americas. The "New World" vultures were widespread ...
). It breeds in forest and savannah across
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
, usually near water, its range coinciding with that of the
oil An oil is any nonpolar chemical substance that is composed primarily of hydrocarbons and is hydrophobic (does not mix with water) & lipophilic (mixes with other oils). Oils are usually flammable and surface active. Most oils are unsaturated ...
and
Raffia palm Raffia palms (''Raphia'') are a genus of about twenty species of palms native to tropical regions of Africa, and especially Madagascar, with one species (''R. taedigera'') also occurring in Central and South America. ''R. taedigera'' is the sour ...
s. It is quite approachable, like many African vultures, and can be seen near habitation, even on large hotel lawns in the tourist areas of countries such as
the Gambia The Gambia,, ff, Gammbi, ar, غامبيا officially the Republic of The Gambia, is a country in West Africa. It is the smallest country within mainland AfricaHoare, Ben. (2002) ''The Kingfisher A-Z Encyclopedia'', Kingfisher Publicatio ...
.


Taxonomy

The palm-nut vulture was formally described in 1788 by the German naturalist
Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich GmelinFerdinand Christoph Oetinger , academic_advisors = , doctora ...
in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
's ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
''. He placed it with the eagles, hawks and falcons in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
'' Falco'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Falco angolensis''. Gmelin based his description on the "Angola vulture" that had been described in 1781 by the English ornithologist John Latham in his multi-volume work ''A General Synopsis of Birds''. The
Leverian Collection The Leverian collection was a natural history and ethnographic collection assembled by Ashton Lever. It was noted for the content it acquired from the voyages of Captain James Cook. For three decades it was displayed in London, being broken up ...
in London included two specimens that had been collected in Angola. The palm-nut vulture is now placed in the genus ''Gypohierax'' that was introduced for the species in 1836 by the German naturalist
Eduard Rüppell Wilhelm Peter Eduard Simon Rüppell (20 November 1794 – 10 December 1884) was a German Natural history, naturalist and List of explorers, explorer. Rüppell is occasionally transliterated to "Rueppell" for the English alphabet, due to german ort ...
. The genus name combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''gups'' meaning "vulture" with ''hierax'' meaning "hawk". The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
: no
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised.


Description

This is a nearly unmistakable bird as an adult. At in weight, in length and with a wingspan of , this is the smallest
Old World vulture Old World vultures are vultures that are found in the Old World, i.e. the continents of Europe, Asia and Africa, and which belong to the family Accipitridae, which also includes eagles, buzzards, kites, and hawks. Old World vultures are not clos ...
. Its plumage is all white except for black areas in its wings and tail. It has a red patch around each eye. The juvenile, which takes 3–4 years to mature, is brown with yellow eye-patches. In flight, this species resembles an eagle more than a typical vulture, and it can sustain flapping flight, so it does not depend on thermals. With its extensive white plumage, and black wing- and tail-feathers, the adult palm-nut vulture can be crudely mistaken for both the African fish-eagle and the
Egyptian vulture The Egyptian vulture (''Neophron percnopterus''), also called the white scavenger vulture or pharaoh's chicken, is a small Old World vulture and the only member of the genus ''Neophron''. It is widely distributed from the Iberian Peninsula and ...
, but clearly lacks the chestnut body of the former and the white tail of the latter The sexes are identical in appearance, with the female being the same size as the male. Juveniles on the other hand are predominately brown with partially black wings and take a lengthy three to four years to make the transition into the adult plumage


Distribution and habitat

Palm-nut vultures are found throughout most of the coastal areas of the African continent from The Gambia to Kenya and as far South as South Africa. The total African population is estimated to be 80 000 pairs. There are approximately 40 birds in
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
. The only Southern African subregions to have resident breeding pairs of palm-nut vultures are South Africa and Mozambique. The breeding distribution of the palm-nut vulture during the 1970s census period was centred on the
Raffia palm Raffia palms (''Raphia'') are a genus of about twenty species of palms native to tropical regions of Africa, and especially Madagascar, with one species (''R. taedigera'') also occurring in Central and South America. ''R. taedigera'' is the sour ...
groves of the
Kosi Bay Kosi Bay is a series of four interlinked lakes in the Maputaland area of KwaZulu-Natal, South Africa. Ecology The lakes form part of the iSimangaliso Wetland Park, a UNESCO World Heritage Site. The closest town is Manguzi, some away from it. K ...
system and
Mtunzini Mtunzini ( Zulu: ''eMthunzini'', from ''umthunzi'' meaning "a place in the shade") is a small coastal town that is situated almost exactly halfway along KwaZulu-Natal's coastline in South Africa approximately 140 km north of Durban. In 2011, ...
. Its distribution is linked to the presence of the Raffia palm ''
Raphia australis ''Raphia australis'', the giant palm or rafia, is a species of raffia palm in the family Arecaceae. It is found around Kosi Bay in southern Mozambique and northeastern KwaZulu Natal in South Africa. It is threatened by habitat loss caused by drai ...
'' at all permanently occupied sites, and the existence of this species at Mtunzini is entirely due to the artificial cultivation of Raffia palms. Currently there are 7 known nesting sites in South Africa and a total of 40 individual birds. As the name suggests, the distribution of the palm-nut vulture closely tracks that of oil (''
Elaeis guineensis ''Elaeis guineensis'' is a species of palm commonly just called oil palm but also sometimes African oil palm or macaw-fat. It is the principal source of palm oil. It is native to west and southwest Africa, specifically the area between Angola an ...
'') or raffia ('' Raphia'' sp.) palms. Consequently, it is most common in coastal forests and mangrove swamps below , but also occurs in wet savannas.


Behaviour and ecology


Food and feeding

Unusually for birds of prey, the palm-nut vulture feeds mainly on the fleshy fruit-husks of the oil palm and on the palm-fruits of Raffia palm. These fruits make up over 60% of the adult bird's diet and over 90% of the juvenile bird's diet. It has also been recorded to feed on crabs (both freshwater and marine),
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant taxon, extant species of molluscs are recognized. The number of fossil sp ...
, frogs, fish, locusts, small mammals, even
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s’ eggs and hatchlings, and it has been known to occasionally attack domestic poultry and feed on
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
.


Breeding

Breeding pairs construct large stick nests high up in tall trees and will often exhibit a strong attachment to the nest site. They may stay at the nesting site for an entire year. Where Raphia Palms are present, breeding pairs will build a nest at the base of the palm fronds. At the beginning of the breeding season, pairs soar together in an aerial display of rolling and diving, much more acrobatic than most vultures. During each breeding cycle, a single, white and brown egg is laid, which is incubated by both sexes, over a period of four to six weeks. Normally around 85 to 90 days after hatching, the young brown chicks will fledge.


Conservation status

This species is widespread throughout much of Africa, overall fairly common and with a stable population. It is rarer and much more localized in South Africa, although not considered to be under any form of immediate threat in that country. That being said, the threats to this species in South Africa are not well understood. The low population size makes the species vulnerable to stochastic events. In Mozambique Parker (1999) observed that cutting down of coastal forest was threatening breeding habitat. The main threat to this species in South Africa is habitat loss. Open cast sand dune mining and urban expansion have reduced suitable habitats. The cultivation of Raphia palms for its ornamental value is currently increasing providing food and nesting sites. There is also a large portion of its habitat protected by the
Isimangaliso Wetland Park iSimangaliso Wetland Park (previously known as the Greater St. Lucia Wetland Park) is situated on the east coast of KwaZulu-Natal, South Africa, about 235 kilometres north of Durban by road. It is South Africa's third-largest protected area, ...
. There are no current species specific conservation initiatives as this species is the only vulture species in South Africa where the population size is increasing. The importance of maintaining this is that the South African satellite population in producing dispersing birds that are contributing to maintaining the Mozambique population (and hence the larger metapopulation). Laika ac Selous (9859653096).jpg, Immature bird in
Selous Game Reserve The Selous Game Reserve, now renamed Nyerere National Park, is a protected area in southern Tanzania. It covers a total area of and has additional buffer zones. It was designated a UNESCO World Heritage Site in 1982 due to its wildlife diversity ...
, Tanzania Palm nut vulture (Gypohierax angolensis) in flight, Semliki Wildlife Reserve.jpg, Adult in flight in
Semliki Wildlife Reserve The Semliki Wildlife Reserve is a conservation protected area in the Western Region of Uganda with headquarters at Karugutu in Ntoroko District. Geography Semliki Wildlife Reserve is located in Kabarole and Ntoroko districts, in the Toro sub ...
, Uganda Palm nut vulture (Gypohierax angolensis) on nest.jpg, Adult on nest at
Kazinga Channel The Kazinga Channel in Uganda is a wide, long natural channel that links Lake Edward and Lake George (Uganda), Lake George, and a dominant feature of Queen Elizabeth National Park. The channel attracts a varied range of animals and birds, with ...
, Uganda


References


Further reading

*


External links

* Palmnut Vulture
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q732523
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the o ...
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the o ...
Birds of the Gulf of Guinea Birds of prey of Sub-Saharan Africa
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the o ...
palm-nut vulture The palm-nut vulture (''Gypohierax angolensis'') or vulturine fish eagle, is a large bird of prey in the family Accipitridae (which also includes many other diurnal raptors such as kites, buzzards and harriers, vultures, and eagles). It is the o ...