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anatomy Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old scien ...
of
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s, the paleocortex, or paleopallium, is a region within the
telencephalon The cerebrum (: cerebra), telencephalon or endbrain is the largest part of the brain, containing the cerebral cortex (of the two cerebral hemispheres) as well as several subcortical structures, including the hippocampus, basal ganglia, and olf ...
in the vertebrate
brain The brain is an organ (biology), organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It consists of nervous tissue and is typically located in the head (cephalization), usually near organs for ...
. This type of cortical tissue consists of three cortical laminae (layers of neuronal cell bodies). In comparison, the
neocortex The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, ...
has six layers and the
archicortex The archicortex, or archipallium, is the phylogenetically second oldest region of the brain's cerebral cortex (the oldest is the paleocortex). In older species, such as fish, the archipallium makes up most of the cerebrum. Amphibians develop an a ...
has three or four layers. Because the number of laminae that compose a type of cortical tissue seems to be directly proportional to both the information-processing capabilities of that tissue and its phylogenetic age, paleocortex is thought to be an intermediate between the
archicortex The archicortex, or archipallium, is the phylogenetically second oldest region of the brain's cerebral cortex (the oldest is the paleocortex). In older species, such as fish, the archipallium makes up most of the cerebrum. Amphibians develop an a ...
(or archipallium) and the
neocortex The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, ...
(or neopallium) in both aspects. The paleocortex (or paleopallium) and the
archicortex The archicortex, or archipallium, is the phylogenetically second oldest region of the brain's cerebral cortex (the oldest is the paleocortex). In older species, such as fish, the archipallium makes up most of the cerebrum. Amphibians develop an a ...
(or the archipallium) of the
cerebral cortex The cerebral cortex, also known as the cerebral mantle, is the outer layer of neural tissue of the cerebrum of the brain in humans and other mammals. It is the largest site of Neuron, neural integration in the central nervous system, and plays ...
together constitute the mammalian
allocortex The allocortex (from Latin allo-, meaning other, and cortex, meaning bark or crust), or heterogenetic cortex, is one of the two types of cerebral cortex in the brain, together with the neocortex. In the human brain, the allocortex is the much sm ...
or the heterogenetic cortex. The distinction for what is called
neocortex The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, ...
or isocortex, which comprises most of the human brain (about 90%), is made from the number of cellular layers that the structure comprises. Neocortical tissue comprises six distinct cell layers, not seen in paleocortical tissues either in adult or developing stage. In humans the paleocortex is exemplified in the olfactory cortex. For most vertebrates, the
olfactory bulb The olfactory bulb (Latin: ''bulbus olfactorius'') is a neural structure of the vertebrate forebrain involved in olfaction, the sense of smell. It sends olfactory information to be further processed in the amygdala, the orbitofrontal cortex (OF ...
is the main feature of the paleocortex, even though the division is virtually unused outside of a mammalian context.


Locations

Paleocortex is present in the
parahippocampal gyrus The parahippocampal gyrus (or hippocampal gyrus') is a grey matter cortical region, a gyrus of the brain that surrounds the hippocampus and is part of the limbic system. The region plays an important role in memory encoding and retrieval. It ha ...
,
olfactory bulb The olfactory bulb (Latin: ''bulbus olfactorius'') is a neural structure of the vertebrate forebrain involved in olfaction, the sense of smell. It sends olfactory information to be further processed in the amygdala, the orbitofrontal cortex (OF ...
, accessory olfactory bulb,
olfactory tubercle The olfactory tubercle (OT), also known as the tuberculum olfactorium, is a multi-sensory processing center that is contained within the olfactory cortex and ventral striatum and plays a role in reward cognition. The OT has also been shown to ...
,
piriform cortex The piriform cortex, or pyriform cortex, is a region in the brain, part of the rhinencephalon situated in the cerebrum. The function of the piriform cortex relates to the sense of smell. Structure The piriform cortex is part of the rhinencephal ...
, periamygdalar area,
anterior olfactory nucleus The anterior olfactory nucleus (AON) also called the anterior olfactory cortex, is a major early processing area for olfaction located behind the olfactory bulb, and in the olfactory tract (olfactory peduncle). The AON connects the olfactory pro ...
,
anterior perforated substance The anterior perforated substance is a part of the brain. It is bilateral. It is irregular and quadrilateral. It lies in front of the optic tract and behind the olfactory trigone. Structure The anterior perforated substance is bilateral. It ...
, and prepyriform area.


Parts of the paleocortex

*
Olfactory bulb The olfactory bulb (Latin: ''bulbus olfactorius'') is a neural structure of the vertebrate forebrain involved in olfaction, the sense of smell. It sends olfactory information to be further processed in the amygdala, the orbitofrontal cortex (OF ...
*
Piriform cortex The piriform cortex, or pyriform cortex, is a region in the brain, part of the rhinencephalon situated in the cerebrum. The function of the piriform cortex relates to the sense of smell. Structure The piriform cortex is part of the rhinencephal ...
*
Anterior olfactory nucleus The anterior olfactory nucleus (AON) also called the anterior olfactory cortex, is a major early processing area for olfaction located behind the olfactory bulb, and in the olfactory tract (olfactory peduncle). The AON connects the olfactory pro ...
*
Olfactory tract The olfactory tract (olfactory peduncle or olfactory stalk) is a bilateral bundle of afferent nerve fibers from the mitral and tufted cells of the olfactory bulb that connects to several target regions in the brain, including the piriform cort ...
*
Anterior commissure The anterior commissure (also known as the precommissure) is a white matter nerve tract, tract (a bundle of axons) connecting the two temporal lobes of the cerebral hemispheres across the midline, and placed in front of the columns of the Fornix o ...
* Uncus


See also

* Paralimbic cortex *
List of regions in the human brain The human brain anatomical regions are ordered following standard neuroanatomy hierarchies. Functional, connective, and developmental regions are listed in parentheses where appropriate. Hindbrain (rhombencephalon) Myelencephalon * M ...


References

Parent, A. (1996). Carpenter's human neuroanatomy (9th ed.). Philadelphia: Williams &Wilkins. {{Authority control Cerebrum Hippocampus (brain) Cerebral cortex