Pale-billed Woodpecker
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The pale-billed woodpecker (''Campephilus guatemalensis'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in subfamily Picinae of the
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. ...
family Picidae. It is found from
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
to
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
.


Taxonomy and systematics

The pale-billed woodpecker has these three subspecies: *''C. g. regius'' Reichenbach, 1854 *''C. g. nelsoni'' ( Ridgway, 1911) *''C. g. guatemalensis'' ( Hartlaub, 1844)


Description

The pale-billed woodpecker is long. The
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''C. g. guatemalensis'' weighs and subspecies ''C. g. regius'' weighs . Both sexes of the nominate subspecies have black upperparts from nape to rump. They have vertical white or buffy white stripes on the sides of their neck that continue onto their back and nearly or fully form a "V". Their tail is dull brown with dull olive-brown on the underside of the outer feathers. Their wings are black except for the outer
primaries Primary elections, or direct primary are a voting process by which voters can indicate their preference for their party's candidate, or a candidate in general, in an upcoming general election, local election, or by-election. Depending on the c ...
, whose upper surface has dull brown outer webs above and are greenish olive-yellow below. Their throat and upper breast are black and the rest of their underparts light yellowish buff with narrow black bars. Their bill is a long ivory yellow chisel, their iris light cream-buff, and their legs greenish gray. Adult males have a red head with a bushy crest and sometimes a grayish brown spot on the ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
. Adult females have a black forecrown, center of the crown, foreneck, and throat. Juveniles resemble adult females but with a darker bill and are mostly dull black on the side of their head.Schulenberg, T. S. (2020). Pale-billed Woodpecker (''Campephilus guatemalensis''), version 1.0. In Birds of the World (T. S. Schulenberg, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.pabwoo1.01 retrieved February 8, 2023 Subspecies ''C. g. nelsoni'' is slightly smaller than the nominate. Its black parts are browner, the upperparts' stripes whiter, and there is often some brown and black barring on the lower back and rump. Subspecies ''C. g. regius'' is larger than the nominate but otherwise similar.


Distribution and habitat

The subspecies of the pale-billed woodpecker are found thus: *''C. g. regius'', eastern Mexico between
Tamaulipas Tamaulipas (), officially the Free and Sovereign State of Tamaulipas ( es, Estado Libre y Soberano de Tamaulipas), is a state in the northeast region of Mexico; one of the 31 states which, along with Mexico City, comprise the 32 Federal Entiti ...
and
Veracruz Veracruz (), formally Veracruz de Ignacio de la Llave (), officially the Free and Sovereign State of Veracruz de Ignacio de la Llave ( es, Estado Libre y Soberano de Veracruz de Ignacio de la Llave), is one of the 31 states which, along with Me ...
*''C. g. nelsoni'', western Mexico between
Sonora Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is d ...
and
Oaxaca Oaxaca ( , also , , from nci, Huāxyacac ), officially the Free and Sovereign State of Oaxaca ( es, Estado Libre y Soberano de Oaxaca), is one of the 32 states that compose the political divisions of Mexico, Federative Entities of Mexico. It is ...
*''C. g. guatemalensis'', from Veracruz,
Chiapas Chiapas (; Tzotzil language, Tzotzil and Tzeltal language, Tzeltal: ''Chyapas'' ), officially the Free and Sovereign State of Chiapas ( es, Estado Libre y Soberano de Chiapas), is one of the states that make up the Political divisions of Mexico, ...
, and the
Yucatán Peninsula The Yucatán Peninsula (, also , ; es, Península de Yucatán ) is a large peninsula in southeastern Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north ...
south through Belize, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica into western Panama The pale-billed woodpecker inhabits the interior and edges of a variety of forested landscapes. The most common of these is lowland tropical
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zone ...
. On the Pacific slope, which is drier than the Caribbean slope, it also occurs in tropical deciduous forest and mangroves. It occurs locally in montane evergreen forest and pine-oak forest in Honduras. It shuns extensive deforested areas. In elevation it ranges from sea level to in Mexico, to in Honduras, to on Costa Rica's Pacific side but only to on the Caribbean side, and to in Panama.


Behavior


Movement

The pale-billed woodpecker is a year-round resident throughout its range.


Feeding

The pale-billed woodpecker forages mostly in the forest's mid- to upper levels but will feed on the ground at stumps and fallen logs. It generally seeks its prey on tree trunks and large limbs, excavating in decayed wood and scaling bark from dead. It forages singly or in pairs, though pairs may be separated by several meters. It has not been seen to join
mixed species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
s. Its diet is not known in detail but is believed to be mostly large larvae of wood-boring beetles; the diet also includes termites and some small fruits.


Breeding

The pale-billed woodpecker's breeding season has not been fully defined but appears to vary with latitude. In southern Mexico it is between December and June, and in Costa Rica it is between August and September. Both sexes excavate the nest cavity in a large, often dead, tree; reported nests have been above the ground. The clutch size is two eggs and both parents incubate. The incubation period, time to fledging, and details of parental care are not known.


Vocal and non-vocal sounds

The pale-billed woodpecker's characteristic call is "a variable, nasal chatter or sputter", described more fully as "a nasal (like a sqeeze toy) ''kip kip kip kip-hey'eh'eh''." It also makes "low whining notes", a "low whining or moaning call", and a "loud 'bleating' call". Its drum is "a loud, rapid, double rap" and, much more rarely, "a resonant series of up to seven rapid raps".


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the pale-billed woodpecker as being of Least Concern. It has a very large range and an estimated population of at least 50,000 mature individuals, though the latter is believed to be decreasing. No immediate threats have been identified. "This woodpecker requires large trees for foraging, however, and it disappears from areas that are deforested or heavily cut over." Populations in Mexico are listed as "subject to special protection" by the Mexican Official Norm NOM-059-SEMARNAT-2010.


References


Further reading

* *


External links

* {{Taxonbar, from=Q1035440 Pale-billed woodpecker Birds of Belize Birds of Central America Birds of Costa Rica Birds of the Yucatán Peninsula pale-billed woodpecker