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In
anatomy Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
, the palatine bones () are two
irregular bone The irregular bones are bones which, from their peculiar form, cannot be grouped as long, short, flat or sesamoid bones. Irregular bones serve various purposes in the body, such as protection of nervous tissue (such as the vertebrae protect th ...
s of the facial skeleton in many animal species, located above the
uvula The palatine uvula, usually referred to as simply the uvula, is a conic projection from the back edge of the middle of the soft palate, composed of connective tissue containing a number of racemose glands, and some muscular fibers. It also conta ...
in the
throat In vertebrate anatomy, the throat is the front part of the neck, internally positioned in front of the vertebrae. It contains the pharynx and larynx. An important section of it is the epiglottis, separating the esophagus from the trachea (windpip ...
. Together with the
maxillae The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
, they comprise the
hard palate The hard palate is a thin horizontal bony plate made up of two bones of the facial skeleton, located in the roof of the mouth. The bones are the palatine process of the maxilla and the horizontal plate of palatine bone. The hard palate spans t ...
. (''Palate'' is derived from the Latin ''palatum''.)


Structure

The palatine bones are situated at the back of the
nasal cavity The nasal cavity is a large, air-filled space above and behind the human nose, nose in the middle of the face. The nasal septum divides the cavity into two cavities, also known as fossae. Each cavity is the continuation of one of the two nostrils. ...
between the
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
and the pterygoid process of the sphenoid bone. They contribute to the walls of three cavities: the floor and lateral walls of the nasal cavity, the roof of the mouth, and the floor of the orbits. They help to form the pterygopalatine and
pterygoid fossa The pterygoid fossa is an anatomical term for the fossa formed by the divergence of the lateral pterygoid plate and the medial pterygoid plate of the sphenoid bone. Structure The lateral and medial pterygoid plates (of the pterygoid process of ...
e, and the
inferior orbital fissure The inferior orbital fissure is formed by the sphenoid bone and the maxilla. It is located posteriorly along the boundary of the floor and lateral wall of the orbit. It transmits a number of structures, including: * the zygomatic branch of the m ...
s. Each palatine bone somewhat resembles the letter L, and consists of a
horizontal plate The horizontal plate of palatine bone is a quadrilateral part of the palatine bone, and has two surfaces and four borders. Surfaces The superior surface, concave from side to side, forms the back part of the floor of the nasal cavity. The inf ...
, a perpendicular plate, and three projecting processes—the pyramidal process, which is directed backward and lateral from the junction of the two parts, and the orbital and sphenoidal processes, which surmount the vertical part, and are separated by a deep notch, the sphenopalatine notch. The two plates form the posterior part of the hard palate and the floor of the nasal cavity; anteriorly, they join with the maxillae. The two horizontal plates articulate with each other at the posterior part of the median palatine suture and more anteriorly with the maxillae at the transverse palatine suture. The human palatine articulates with six bones: the sphenoid,
ethmoid The ethmoid bone (; from grc, ἡθμός, hēthmós, sieve) is an unpaired bone in the skull that separates the nasal cavity from the brain. It is located at the roof of the nose, between the two orbits. The cubical bone is lightweight due to a ...
,
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
,
inferior nasal concha The inferior nasal concha (inferior turbinated bone or inferior turbinal/turbinate) is one of the three paired nasal conchae in the nose. It extends horizontally along the lateral wall of the nasal cavity and consists of a lamina of spongy bone, c ...
,
vomer The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right max ...
and opposite palatine. There are two important foramina in the palatine bones that transmit nerves and blood vessels to this region: the greater and lesser palatine. The larger
greater palatine foramen At either posterior angle of the hard palate is the greater palatine foramen, for the transmission of the descending palatine vessels and greater palatine nerve; and running anteriorly (forward) and medially (towards the center-line) from it is a g ...
is located in the posterolateral region of each of the palatine bones, usually at the apex of the maxillary third molar. The greater palatine foramen transmits the greater palatine nerve and blood vessels. A smaller opening nearby, the
lesser palatine foramen Behind the greater palatine foramen is the pyramidal process of the palatine bone, perforated by one or more lesser palatine foramina which carry the lesser palatine nerve, and marked by the commencement of a transverse ridge, for the attachment o ...
, transmits the lesser palatine nerve and blood vessels to the soft palate and tonsils. Both foramina are openings of the pterygopalatine canal that carries the descending palatine nerves and blood vessels from the pterygopalatine fossa to the palate.


Function

The
sphenopalatine foramen The sphenopalatine foramen is a Foramina of the skull, foramen in the skull that connects the nasal cavity with the pterygopalatine fossa. Structure The processes of the superior border of the palatine bone are separated by the ''sphenopalatine no ...
is the opening between the sphenoid bone and orbital processes of the palatine bone; it opens into the nasal cavity and gives passage to branches from the pterygopalatine ganglion and the
sphenopalatine artery The sphenopalatine artery (nasopalatine artery) is an artery of the head, commonly known as the artery of epistaxis. Course The sphenopalatine artery is a branch of the maxillary artery which passes through the sphenopalatine foramen into the cavi ...
from the maxillary artery.


Other animals

In
bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...
the palatine bone consists of the perpendicular plate only, lying on the inner edge of the maxilla. The lower surface of the bone may bear several teeth, forming a second row behind those of the maxilla; in many cases, these are actually larger than the maxillary teeth. Although a similar pattern was present in primitive
tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct theraps ...
s, the palatine bone is reduced in most living
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
s, forming, in
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s and
salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All t ...
s, only a narrow bar between the vomer and maxilla. Early fossil
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s retained the arrangement seen in more primitive vertebrates, but in
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s, the lower surface of the palatine became folded over during evolution, forming the horizontal plate, and meeting in the midline of the mouth. This forms the rear of the
hard palate The hard palate is a thin horizontal bony plate made up of two bones of the facial skeleton, located in the roof of the mouth. The bones are the palatine process of the maxilla and the horizontal plate of palatine bone. The hard palate spans t ...
, separating the oral and nasal cavities, and making it easier to breathe while eating. A
parallel Parallel is a geometric term of location which may refer to: Computing * Parallel algorithm * Parallel computing * Parallel metaheuristic * Parallel (software), a UNIX utility for running programs in parallel * Parallel Sysplex, a cluster of ...
development has occurred to varying degrees in many living reptiles, reaching its greatest extent in
crocodilian Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
s. In
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s, the palatine bones remain separate, long the sides of the rear part of the upper jaw, and typically have a mobile articulation with the
cranium The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
. There are numerous variations amongst mammals, amphibians and other species. For example, the palatine bone in many amphibians such as the
rough-skinned newt The rough-skinned newt or roughskin newt (''Taricha granulosa'') is a North American newt known for the strong toxin exuded from its skin. Appearance A stocky newt with rounded snout, it ranges from light brown to olive or brownish-black on ...
manifests as a distinct V-shaped structure. In the case of cat species, the horizontal and a vertical elements join at a forty five degree angle.


Additional images

File:Orbital_bones.png, The seven bones which articulate to form the orbit. File:Gray167.png, Articulation of left palatine bone with maxilla. File:Gray187.png, Base of skull. Inferior surface. File:Gray192.png, Medial wall of left orbit. File:Gray196.png, Roof, floor, and lateral wall of left nasal cavity. File:Gray168.png, Medial aspect of left palatine bone. File:Gray169.png, Posterior aspect of left palatine bone.


See also

**
Nasopalatine nerve The nasopalatine nerve (long sphenopalatine nerve) is a nerve of the head. It is a branch of the pterygopalatine ganglion, a continuation from the maxillary nerve (V2). It supplies parts of the palate and nasal septum. Structure The nasopalati ...
** Superior nasal nerve * Bone terminology *
Terms for anatomical location Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...


References


External links

{{Authority control Bones of the head and neck Irregular bones