Palaeoscincus
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''Palaeoscincus'' (meaning "ancient
skink Skinks are lizards belonging to the family Scincidae, a family in the infraorder Scincomorpha. With more than 1,500 described species across 100 different taxonomic genera, the family Scincidae is one of the most diverse families of lizards. Ski ...
" from the Greek παλαιός and σκίγγος) is a
dubious Doubt is a mental state in which the mind remains suspended between two or more contradictory propositions, unable to be certain of any of them. Doubt on an emotional level is indecision between belief and disbelief. It may involve uncertainty, ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
ankylosauria Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
based on
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
from the mid-late
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
-age
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Judith River Formation The Judith River Formation is a fossil-bearing geologic formation in Montana, and is part of the Judith River Group. It dates to the Late Cretaceous, between 79 and 75.3 million years ago, corresponding to the "Judithian" land vertebrate age. It ...
of
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
. Like several other dinosaur genera named by
Joseph Leidy Joseph Mellick Leidy (September 9, 1823 – April 30, 1891) was an American paleontologist, parasitologist and anatomist. Leidy was professor of anatomy at the University of Pennsylvania, later was a professor of natural history at Swarthmore ...
(''
Deinodon ''Deinodon'' (Greek for "terrible tooth") is a dubious tyrannosaurid dinosaur genus containing a single species, ''Deinodon horridus''. ''D. horridus'' is known only from a set of teeth found in the Late Cretaceous Judith River Formation of Monta ...
'', '' Thespesius'', and ''
Trachodon ''Trachodon'' (meaning "rough tooth") is a dubious genus of hadrosaurid dinosaur based on teeth from the Campanian-age Upper Cretaceous Judith River Formation of Montana, U.S.Leidy, J. (1856). "Notice of remains of extinct reptiles and fishes, d ...
''), it is an historically important genus with a convoluted taxonomy that has been all but abandoned by modern dinosaur
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
s. Because of its wide use in the early 20th century, it was somewhat well known to the general public, often through illustrations of an animal with the
armor Armour (British English) or armor (American English; see spelling differences) is a covering used to protect an object, individual, or vehicle from physical injury or damage, especially direct contact weapons or projectiles during combat, or fr ...
of ''
Edmontonia ''Edmontonia'' is a genus of panoplosaurin nodosaurid dinosaur from the Late Cretaceous Period. It is part of the Nodosauridae, a family within Ankylosauria. It is named after the Edmonton Formation (now the Horseshoe Canyon Formation in Canada) ...
'' and the tail
club Club may refer to: Arts, entertainment, and media * ''Club'' (magazine) * Club, a ''Yie Ar Kung-Fu'' character * Clubs (suit), a suit of playing cards * Club music * "Club", by Kelsea Ballerini from the album ''kelsea'' Brands and enterprises ...
of an
ankylosaurid Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...
.


Reassigned species

Seven species have been referred to this genus over the years, six of which have since been reassigned to other genera: *''Palaeoscincus africanus'', named by
Robert Broom Robert Broom FRS FRSE (30 November 1866 6 April 1951) was a British- South African doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University of Glasgow. From 1903 to 1910, he ...
in 1910/1912, a partial
jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...
from the
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
-
Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretaceou ...
-age
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
-
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...
Kirkwood Formation The Kirkwood Formation is a geological Geological formation, formation found in the Eastern and Western Cape provinces in South Africa. It is one of the four formations found within the Uitenhage Group of the Algoa Basin – its type locality – ...
of
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, now known as the
stegosaurid Stegosauridae is a family of thyreophoran dinosaurs (armoured dinosaurs) within the suborder Stegosauria. The clade is defined as all species of dinosaurs more closely related to ''Stegosaurus'' than ''Huayangosaurus''.David B. Weishampel, Peter ...
''
Paranthodon ''Paranthodon'' ( ) is a genus of stegosaurian dinosaur that lived in what is now South Africa during the Early Cretaceous, between 139 and 131 million years ago. Discovered in 1845, it was one of the first stegosaurians found. Its only rem ...
''; *''Palaeoscincus asper'', "the rough one", a dubious tooth
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
from the late Campanian-age Upper Cretaceous
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
of
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, named by
Lawrence Morris Lambe Lawrence Morris Lambe (August 27, 1863 – March 12, 1919) was a Canadian geologist, palaeontologist, and ecologist from the Geological Survey of Canada (GSC). His published work, describing the diverse and plentiful dinosaur discoveries from th ...
in 1902, based on a single tooth, specimen NMC 1349 now referred to ''
Euoplocephalus ''Euoplocephalus'' ( ) is a genus of very large, herbivorous ankylosaurid dinosaurs, living during the Late Cretaceous of Canada. It has only one named species, ''Euoplocephalus tutus''. The first fossil of ''Euoplocephalus'' was found in 1897 i ...
'';Coombs Jr., W.P. (1990). Teeth and taxonomy in ankylosaurs. In: Carpenter, K., and Currie, P.J. (eds.). ''Dinosaur Systematics: Approaches and Perspectives''. Cambridge University Press:Cambridge, 269-279. *''Palaeoscincus costatus'', "the ribbed one", the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
named by Leidy in 1856, known from a single tooth, specimen ANSP 9263 found by
Ferdinand Vandeveer Hayden Ferdinand Vandeveer Hayden (September 7, 1829 – December 22, 1887) was an American geologist noted for his pioneering surveying expeditions of the Rocky Mountains in the late 19th century. He was also a physician who served with the Union Ar ...
near Fort Benton. It was the first ankylosaurian species to be named based on American material; it is now considered an ankylosaurian of unknown affinities. *''Palaeoscincus latus'', "the wide one" named by
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among h ...
in 1892, from the late Maastrichtian-age Upper Cretaceous
Lance Formation The Lance (Creek) Formation is a division of Late Cretaceous (dating to about 69 - 66 Ma) rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the late ...
of
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
, also based on a single tooth, specimen YPM 4810 found in Niobrara County,
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
, now believed to have come from a
pachycephalosaur Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachyc ...
id; *"P. magoder", a ''
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
'' name from a
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
l list by Karl ("Charles") L. Henning, the result of mistaking the German words ''mag oder'' for a specific name; *''Palaeoscincus rugosidens'', "rough tooth" named by
Charles Whitney Gilmore Charles Whitney Gilmore (March 11, 1874 – September 27, 1945) was an American paleontologist who gained renown in the early 20th century for his work on vertebrate fossils during his career at the United States National Museum (now the N ...
in 1930, the best-known species, a
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
and partial skeleton from the late
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
-age
Two Medicine Formation The Two Medicine Formation is a geological formation, or rock body, in northwestern Montana and southern Alberta that was deposited between and (million years ago), during Campanian (Late Cretaceous) time. It crops out to the east of the Rocky M ...
of
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
, now known as ''Edmontonia rugosidens'', or a separate genus '' Chassternbergia''. It was this species that was portrayed in most restorations of the genus. *''Palaeoscincus tutus'', a renaming of ''Euoplocephalus tutus'' by
Edwin Hennig Edwin Hennig (27 April 1882 – 12 November 1977) was a German paleontologist. Career Edwin Hennig was one of five children of a merchant who died when Hennig was 10 years old. Starting in 1902, Hennig studied natural sciences, anthropolog ...
in 1915. Today, the type species ''P. costatus'' and thereby the genus is considered to be an indeterminate ankylosaurian,Carpenter, K. (2001). Phylogenetic analysis of the Ankylosauria. In: Carpenter, K. (ed.). ''The Armored Dinosaurs''. Indiana University Press:Bloomington, 455-483. Vickaryous, M.K., Maryańska, T., and Weishampel, D.B., (2004). Ankylosauria. In: Weishampel, D.B., Dodson, P., and Osmólska, H. (eds.). ''The Dinosauria (second edition)''. University of California Press:Berkeley, 363-392. perhaps an indeterminate
nodosaurid Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, wer ...
.Ford, T.L. (2000). A review of ankylosaur osteoderms from New Mexico and a preliminary review of ankylosaur armor. In: Lucas, S.G., and Heckert, A.B. (eds.). ''Dinosaurs of New Mexico''. ''New Mexico Museum of Natural History and Science Bulletin'' 17:157-176.


See also

*
Timeline of ankylosaur research This timeline of ankylosaur research is a chronological listing of events in the history of paleontology focused on the ankylosaurs, quadrupedal herbivorous dinosaurs who were protected by a covering bony plates and spikes and sometimes by a club ...


References


External links


Quick taxonomic summary
from the Dinosaur Mailing List {{Taxonbar, from=Q871916 Nodosaurids Late Cretaceous dinosaurs of North America Nomina dubia Fossil taxa described in 1856 Taxa named by Joseph Leidy Paleontology in Montana Campanian genus first appearances Campanian genus extinctions