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The Oligocene ( ) is a geologic
epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided ...
of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from the Ancient Greek (''olígos'', "few") and (''kainós'', "new"), and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', ...
Epoch and is followed by the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s of the Miocene. Major changes during the Oligocene included a global expansion of
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s, and a regression of
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
broad leaf
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s to the equatorial belt. The start of the Oligocene is marked by a notable extinction event called the Grande Coupure; it featured the replacement of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
an fauna with
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
n
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is '' flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. ...
, except for the endemic
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
and
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in ...
families. By contrast, the Oligocene–Miocene boundary is not set at an easily identified worldwide event but rather at regional boundaries between the warmer late Oligocene and the relatively cooler Miocene.


Boundaries and subdivisions

The lower boundary of the Oligocene (its
Global Boundary Stratotype Section and Point A Global Boundary Stratotype Section and Point (GSSP) is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. The effort to define GSSPs is conducted ...
or GSSP) is placed at the last appearance of the foraminiferan genus '' Hantkenina'' in a quarry at Massignano,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
. However, this GSSP has been criticized as excluding the uppermost part of the type Eocene Priabonian Stage and because it is slightly earlier than important climate shifts that form natural markers for the boundary, such as the global oxygen isotope shift marking the expansion of Antarctic glaciation (the Oi1 event). The upper boundary of the Oligocene is defined by its GSSP at Carrosio,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
, which coincides with the first appearance of the foraminiferan '' Paragloborotalia kugleri'' and with the base of magnetic polarity chronozone C6Cn.2n. Oligocene faunal stages from youngest to oldest are:


Tectonics and paleogeography

During the Oligocene Epoch, the continents continued to
drift Drift or Drifts may refer to: Geography * Drift or ford (crossing) of a river * Drift, Kentucky, unincorporated community in the United States * In Cornwall, England: ** Drift, Cornwall, village ** Drift Reservoir, associated with the village ...
toward their present positions.
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
became more isolated as deep ocean channels were established between Antarctica and
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
. Australia had been very slowly rifting away from West Antarctica since the Jurassic, but the exact timing of the establishment of ocean channels between the two continents remains uncertain. However, one estimate is that a deep channel was in place between the two continents by the end of the early Oligocene. The timing of the formation of the Drake Passage between South America and Antarctica is also uncertain, with estimates ranging from 49 to 17 mya (early Eocene to Miocene), but oceanic circulation through the Drake Passage may also have been in place by the end of the early Oligocene. This may have been interrupted by a temporary constriction of the Drake Passage from sometime in the middle to late Oligocene (29 to 22 mya) to the middle Miocene (15 mya). The reorganization of the oceanic tectonic plates of the northeastern Pacific, which had begun in the Paleocene, culminated with the arrival of the Murray and Mendocino Fracture Zones at the North American subduction zone in the Oligocene. This initiated strike-slip movement along the
San Andreas Fault The San Andreas Fault is a continental transform fault that extends roughly through California. It forms the tectonic boundary between the Pacific Plate and the North American Plate, and its motion is right-lateral strike-slip (horizontal) ...
and extensional tectonics in the
Basin and Range province The Basin and Range Province is a vast physiographic region covering much of the inland Western United States and northwestern Mexico. It is defined by unique basin and range topography, characterized by abrupt changes in elevation, alternating ...
, ended volcanism south of the Cascades, and produced clockwise rotation of many western North American terranes. The Rocky Mountains were at their peak. A new volcanic arc was established in western North America, far inland from the coast, reaching from central Mexico through the Mogollon-Datil volcanic field to the San Juan volcanic field, then through Utah and Nevada to the ancestral Northern Cascades. Huge ash deposits from these volcanoes created the White River and Arikaree Groups of the High Plains, with their excellent fossil beds. Between 31 and 26 mya, the Ethiopia-Yemen Continental Flood Basalts were emplaced by the East African large igneous province, which also initiated rifting along the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...
and Gulf of Aden. The
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
were rapidly rising in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
as the African plate continued to push north into the
Eurasian plate The Eurasian Plate is a tectonic plate that includes most of the continent of Eurasia (a landmass consisting of the traditional continents of Europe and Asia), with the notable exceptions of the Indian subcontinent, the Arabian subcontinent an ...
, isolating the remnants of the Tethys Sea. Sea levels were lower in the Oligocene than in the early Eocene, exposing large coastal plains in Europe and the Gulf Coast and Atlantic Coast of North America. The
Obik Sea The Turgai Sea, also known as the Turgay Sea, Turgai Strait, Obik Sea, Ural Sea or West Siberian Sea, was a large shallow body of salt water (an epicontinental or epeiric sea) during the Mesozoic through Cenozoic Eras. It extended north of the pres ...
, which had separated Europe from Asia, retreated early in the Oligocene, creating a persistent land connection between the continents. There appears to have been a land bridge in the early Oligocene between North America and Europe, since the
faunas Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is ''funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Z ...
of the two regions are very similar. However, towards the end of the Oligocene, there was a brief marine incursion in Europe. The rise of the Himalayas during the Oligocene remains poorly understood. One recent hypothesis is that a separate microcontinent collided with south Asia in the early Eocene, and India itself did not collide with south Asia until the end of the Oligocene. The
Tibetan Plateau The Tibetan Plateau (, also known as the Qinghai–Tibet Plateau or the Qing–Zang Plateau () or as the Himalayan Plateau in India, is a vast elevated plateau located at the intersection of Central, South and East Asia covering most of the Ti ...
may have reached nearly its present elevation by the late Oligocene. The Andes first became a major mountain chain in the Oligocene, as subduction became more direct into the coastline.


Climate

Climate during the Oligocene reflected a general cooling trend following the Early Eocene Climatic Optimum. This transformed the Earth's climate from a greenhouse to an icehouse climate.


Eocene-Oligocene transition and Oi1 event

The Eocene-Oligocene transition was a major cooling event and reorganization of the biosphere, being part of a broader trend of global cooling lasting from the Bartonian to the Rupelian. The transition is marked by the Oi1 event, an oxygen isotope excursion occurring approximately 33.55 million years ago, during which oxygen isotope ratios decreased by 1.3. About 0.3–0.4 of this is estimated to be due to major expansion of Antarctic ice sheets. The remaining 0.9 to 1.0 was due to about of global cooling. The transition likely took place in three closely spaced steps over the period from 33.8 to 33.5 mya. By the end of the transition, sea levels had dropped by , and ice sheets were 25% greater in extent than in the modern world. The effects of the transition can be seen in the geological record at many locations around the world. Ice volumes rose as temperature and sea levels dropped. Playa lakes of the Tibetan Plateau disappeared at the transition, pointing to cooling and aridification of central Asia. Pollen and spore counts in marine sediments of the Norwegian-Greenland Sea indicate a drop in winter temperatures at high latitudes of about just prior to the Oi1 event. Borehole dating from the Southeast Faroes drift indicates that deep-ocean circulation from the Arctic Ocean to the North Atlantic Ocean began in the early Oligocene. The best terrestrial record of Oligocene climate comes from North America, where temperatures dropped by in the earliest Oligocene. This change is seen from Alaska to the Gulf Coast. Upper Eocene
paleosols In the geosciences, paleosol (''palaeosol'' in Great Britain and Australia) is an ancient soil that formed in the past. The precise definition of the term in geology and paleontology is slightly different from its use in soil science. In ...
reflect annual precipitation of over a meter of rain, but early Oligocene precipitation was less than half this. In central North America, the cooling was by 8.2 ± 3.1 °C over a period of 400,000 years, though there is little indication of significant increase in aridity during this interval. Ice-rafted debris in the Norwegian-Greenland Sea indicated that glaciers had appeared in Greenland by the start of the Oligocene. Continental ice sheets in Antarctica reached sea level during the transition. Glacially rafted debris of early Oligocene age in the Weddell Sea and
Kerguelen Plateau The Kerguelen Plateau (, ), also known as the Kerguelen–Heard Plateau, is an oceanic plateau and a large igneous province (LIP) located on the Antarctic Plate, in the southern Indian Ocean. It is about to the southwest of Australia and is ...
, in combination with Oi1 isotope shift, provides unambiguous evidence of a continental ice sheet on Antarctica by the early Oligocene. The causes of the Eocene-Oligocene transition are not yet fully understood. The timing is wrong for this to be caused either by known impact events or by the volcanic activity on the Ethiopean Plateau. Two other possible drivers of climate change, not mutually exclusive, have been proposed. The first is thermal isolation of the continent of Antarctica by development of the Antarctic Circumpolar Current. Deep sea cores from south of New Zealand suggest that cold deep-sea currents were present by the early Oligocene. However, the timing of this event remains controversial. The other possibility, for which there is considerable evidence, is a drop in atmospheric
carbon dioxide Carbon dioxide ( chemical formula ) is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature. In the air, carbon dioxide is t ...
levels ( pCO2) during the transition. The pCO2 is estimated to have dropped just before the transition, to 760 ppm at the peak of ice sheet growth, then rebounded slightly before resuming a more gradual fall. Climate modeling suggests that glaciation of Antarctica took placed only when pCO2 dropped below a critical threshold value.


Middle Oligocene climate and the Oi2 event

Oligocene climate following the Eocene-Oligocene event is poorly known. There were several pulses of glaciation in middle Oligocene, about the time of the Oi2 oxygen isotope shift. This led to the largest drop of sea level in past 100 million years, by about . This is reflected in a mid-Oligocene incision of continental shelves and unconformities in marine rocks around the world. Some evidence suggests that the climate remained warm at high latitudes even as ice sheets experienced cyclical growth and retreat in response to
orbital forcing Orbital forcing is the effect on climate of slow changes in the tilt of the Earth's axis and shape of the Earth's orbit around the sun (see Milankovitch cycles). These orbital changes modify the total amount of sunlight reaching the Earth by up ...
and other climate drivers. Other evidence indicates significant cooling at high latitudes. Part of the difficulty may be that there were strong regional variations in the response to climate shifts. Evidence of a relatively warm Oligocene suggests an enigmatic climate state, neither hothouse nor icehouse.


Late Oligocene warming

The late Oligocene (26.5 to 24 mya) likely saw a warming trend in spite of low pCO2 levels, though this appears to vary by region. However, Antarctica remained heavily glaciated during this warming period. The late Oligocene warming is discernible in pollen counts from the Tibetan Plateau, which also show that the south Asian monsoon had already developed by the late Oligocene. A deep 400,000-year glaciated Oligocene-Miocene boundary event is recorded at McMurdo Sound and King George Island.


Biosphere

The early Eocene climate was very warm, with crocodilians and temperate plants thriving above the
Arctic Circle The Arctic Circle is one of the two polar circles, and the most northerly of the five major circles of latitude as shown on maps of Earth. Its southern equivalent is the Antarctic Circle. The Arctic Circle marks the southernmost latitude at ...
. The cooling trend that began in the middle Eocene continued into the Oligocene, bringing the poles well below freezing for the first time in the Phanerozoic. The cooling climate, together with the opening of some land bridges and the closing of others, led to a profound reorganization of the biosphere and loss of taxonomic diversity. Land animals and marine organisms reached a Phanerozoic low in diversity by the late Oligocene, and the temperate forests and jungles of the Eocene were replaced by forest and scrubland. The closing of the Tethys Seaway destroyed its tropical biota.


Flora

The Oi1 event of the Eocene-Oligocene transition covered the continent of Antarctica with ice sheets, leaving '' Nothofagus'' and mosses and ferns clinging to life around the periphery of Antarctica in tundra conditions. Angiosperms continued their expansion throughout the world as tropical and sub- tropical forests were replaced by temperate deciduous forests. Open
plain In geography, a plain is a flat expanse of land that generally does not change much in elevation, and is primarily treeless. Plains occur as lowlands along valleys or at the base of mountains, as coastal plains, and as plateaus or uplands ...
s and deserts became more common and
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
es expanded from their water-bank habitat in the Eocene moving out into open tracts. The decline in pCO2 favored C4 photosynthesis, which is found only in angiosperms and is particularly characteristic of grasses. However, even at the end of the period, grass was not quite common enough for modern
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground ...
s. In North America, much of the dense forest was replaced by patchy scrubland with riparian forests. Subtropical species dominated with cashews and
lychee Lychee (US: ; UK: ; ''Litchi chinensis''; ) is a Monotypic taxon, monotypic taxon and the sole member in the genus ''Litchi'' in the Sapindus, soapberry family, ''Sapindaceae''. It is a tropical tree native to Southeast and Southwest China (t ...
trees present, and temperate woody plants such as
rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...
s,
beech Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engl ...
es, and
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanic ...
s were common. The legumes spread, while
sedge The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus '' Carex'' ...
s and
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes exce ...
s continued their ascent.


Fauna

Most extant mammal families had appeared by the end of the Oligocene. These included primitive three-toed horses, rhinoceroses, camels, deer, and peccaries. Carnivores such as dogs, nimravids (ancestor of cats), bears, weasels, and raccoons began to replace the creodonts that had dominated the Paleocene in the Old World. Rodents and rabbits underwent tremendous diversification due to the increase in suitable habitats for ground-dwelling seed eaters, as habitats for squirrel-like nut- and fruit-eaters diminished. The primates, once present in Eurasia, were reduced in range to Africa and South America. Many groups, such as equids, entelodonts, rhinos,
merycoidodont Merycoidodontoidea, sometimes called "oreodonts" or "ruminating hogs", is an extinct Superfamily (taxonomy), superfamily of prehistoric cud-chewing artiodactyls with short faces and fang-like canine teeth. As their name implies, some of the bette ...
s, and camelids, became more able to run during this time, adapting to the plains that were spreading as the Eocene rainforests receded. Brontotheres died out in the Earliest Oligocene, and creodonts died out outside
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
and the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
at the end of the period. Multituberculates, an ancient lineage of primitive mammals that originated back in the
Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
, also became extinct in the Oligocene, aside from the gondwanatheres. The Eocene-Oligocene transition in Europe and Asia has been characterized as the Grande Coupure. The lowering of sea levels closed the Turgai Strait across the Obik Sea, which had previously separated Asia from Europe. This allowed Asian mammals, such as
rhinoceroses A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species ...
and
ruminants Ruminants (suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
, to enter Europe and drive endemic species to extinction. Lesser faunal turnovers occurred simultaneously with the Oi2 event and towards the end of the Oligocene. There was significant diversification of mammals in Eurasia, including the giant indricotheres, that grew up to at the shoulder and weighed up to 20 tons. '' Paraceratherium'' was one of the largest land mammals ever to walk the Earth. However, the indricotheres were an exception to a general tendency for Oligocene mammals to be much smaller than their Eocene counterparts. The earliest deer, giraffes, pigs, and cattle appeared in the mid-Oligocene in Eurasia. The first felid, '' Proailurus'', originated in Asia during the late Oligocene and spread to Europe. There was only limited migration between Asia and North America. The cooling of central North America at the Eocene-Oligocene transition resulted in a large turnover of gastropods, amphibians, and reptiles. Mammals were much less affected. Crocodilians and pond turtles replaced by dry land tortoises. Molluscs shifted to more drought-tolerant forms. The White River Fauna of central North America inhabited a semiarid prairie home and included entelodonts like '' Archaeotherium'', camelids (such as ''
Poebrotherium ''Poebrotherium'' ( ) is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately . Discovery and history ''Poebrotherium'' was first named by scientist ...
''), running rhinoceratoids, three-toed equids (such as ''
Mesohippus ''Mesohippus'' (Greek: / meaning "middle" and / meaning "horse") is an extinct genus of early horse. It lived 37 to 32 million years ago in the Early Oligocene. Like many fossil horses, ''Mesohippus'' was common in North America. Its shoulder hei ...
''), nimravids, protoceratids, and early canids like '' Hesperocyon''. Merycoidodonts, an endemic American group, were very diverse during this time. Australia and South America became geographically isolated and developed their own distinctive endemic fauna. These included the New World and Old World monkeys. The South American continent was home to animals such as pyrotheres and astrapotheres, as well as litopterns and notoungulates.
Sebecosuchian Sebecosuchia is an extinct group of mesoeucrocodylian crocodyliforms that includes the families Sebecidae and Baurusuchidae. The group was long thought to have first appeared in the Late Cretaceous with the baurusuchids and become extinct in t ...
s, terror birds, and carnivorous metatheres, like the borhyaenids remained the dominant predators. Africa was also relative isolated and retained its endemic fauna. These included mastodonts, hyraxes, arsinoitheres, and other archaic forms. Egypt in the Oligocene was an environment of lush forested deltas. At sea, 97% of marine snail species, 89% of clams, and 50% of echinoderms of the Gulf Coast did not survive past the earliest Oligocene. New species evolved, but the overall diversity diminished. Cold-water mollusks migrated around the Pacific Rim from Alaska and Siberia. The marine animals of Oligocene oceans resembled today's fauna, such as the bivalves. Calcareous cirratulids appeared in the Oligocene. The fossil record of marine mammals is a little spotty during this time, and not as well known as the Eocene or Miocene, but some fossils have been found. The baleen whales and toothed whales had just appeared, and their ancestors, the archaeocete
cetacean Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel th ...
s began to decrease in diversity due to their lack of echolocation, which was very useful as the water became colder and cloudier. Other factors to their decline could include climate changes and competition with today's modern cetaceans and the
requiem shark Requiem sharks are sharks of the family Carcharhinidae in the order Carcharhiniformes. They are migratory, live-bearing sharks of warm seas (sometimes of brackish or fresh water) and include such species as the tiger shark, bull shark, le ...
s, which also appeared in this epoch. Early desmostylians, like '' Behemotops'', are known from the Oligocene. Pinnipeds appeared near the end of the epoch from an otter-like ancestor. File:NMNH-USNMV15917Poebrotherium.jpg,
Poebrotherium ''Poebrotherium'' ( ) is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately . Discovery and history ''Poebrotherium'' was first named by scientist ...
File:Merycoidodon Skull Oligocene Left Side.jpg, Merycoidodon File:Hoplophoneus primaevus (fossil false sabertooth cat) (Middle Oligocene; Nebraska, USA) 3 (32791323412).jpg, Hoplophoneus File:Mesohippus barbouri Harvard.jpg,
Mesohippus ''Mesohippus'' (Greek: / meaning "middle" and / meaning "horse") is an extinct genus of early horse. It lived 37 to 32 million years ago in the Early Oligocene. Like many fossil horses, ''Mesohippus'' was common in North America. Its shoulder hei ...
File:Paraceratherium transouralicum skull.jpg, Paraceratherium File:Paleoparadoxia Natural History Museum of Los Angeles County 20110330.jpg, Paleoparadoxia File:Protoceras skeleton.jpg,
Protoceras ''Protoceras'' ('first horns') is an extinct genus of Artiodactyla, of the family Protoceratidae, endemic to North America. It lived from the Oligocene to the Early Miocene 33.3—16.0 Ma, existing for approximately . Morphology ''Protoceras ...
File:Archaeotherium mortoni.JPG, Archaeotherium File:Janjucetus hunderi skull.jpg,
Janjucetus ''Janjucetus'' is an extinct genus of cetacean, and a basal baleen whale (Mysticeti), from the Late Oligocene around 25 million years ago (mya) off southeast Australia, containing one species ''J. hunderi''. Unlike modern mysticetes, it pos ...


Oceans

The Oligocene sees the beginnings of modern ocean circulation, with tectonic shifts causing the opening and closing of ocean gateways. Cooling of the oceans had already commenced by the Eocene/Oligocene boundary, and they continued to cool as the Oligocene progressed. The formation of permanent Antarctic ice sheets during the early Oligocene and possible glacial activity in the Arctic may have influenced this oceanic cooling, though the extent of this influence is still a matter of some significant dispute.


The effects of oceanic gateways on circulation

The opening and closing of ocean gateways: the opening of the Drake Passage; the opening of the Tasmanian Gateway and the closing of the Tethys seaway; along with the final formation of the
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its ...
Faroes The Faroe Islands ( ), or simply the Faroes ( fo, Føroyar ; da, Færøerne ), are a North Atlantic island group and an autonomous territory of the Kingdom of Denmark. They are located north-northwest of Scotland, and about halfway betw ...
Ridge; played vital parts in reshaping oceanic currents during the Oligocene. As the continents shifted to a more modern configuration, so too did ocean circulation.


The Drake Passage

The Drake Passage is located between
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
. Once the Tasmanian Gateway between Australia and Antarctica opened, all that kept Antarctica from being completely isolated by the
Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-smal ...
was its connection to South America. As the South American continent moved north, the Drake Passage opened and enabled the formation of the Antarctic Circumpolar Current (ACC), which would have kept the cold waters of Antarctica circulating around that continent and strengthened the formation of Antarctic Bottom Water (ABW). With the cold water concentrated around Antarctica, sea surface temperatures and, consequently, continental temperatures would have dropped. The onset of Antarctic glaciation occurred during the early Oligocene, and the effect of the Drake Passage opening on this glaciation has been the subject of much research. However, some controversy still exists as to the exact timing of the passage opening, whether it occurred at the start of the Oligocene or nearer the end. Even so, many theories agree that at the Eocene/Oligocene (E/O) boundary, a yet shallow flow existed between South America and Antarctica, permitting the start of an Antarctic Circumpolar Current. Stemming from the issue of when the opening of the Drake Passage took place, is the dispute over how great of an influence the opening of the Drake Passage had on the global climate. While early researchers concluded that the advent of the ACC was highly important, perhaps even the trigger, for Antarctic glaciation and subsequent global cooling, other studies have suggested that the δ18O signature is too strong for glaciation to be the main trigger for cooling. Through study of Pacific Ocean sediments, other researchers have shown that the transition from warm Eocene ocean temperatures to cool Oligocene ocean temperatures took only 300,000 years, which strongly implies that feedbacks and factors other than the ACC were integral to the rapid cooling.


= The late Oligocene opening of the Drake Passage

= The latest hypothesized time for the opening of the Drake Passage is during the early Miocene. Despite the shallow flow between South America and Antarctica, there was not enough of a deep water opening to allow for significant flow to create a true Antarctic Circumpolar Current. If the opening occurred as late as hypothesized, then the Antarctic Circumpolar Current could not have had much of an effect on early Oligocene cooling, as it would not have existed.


= The early Oligocene opening of the Drake Passage

= The earliest hypothesized time for the opening of the Drake Passage is around 30 Ma. One of the possible issues with this timing was the continental debris cluttering up the seaway between the two plates in question. This debris, along with what is known as the Shackleton Fracture Zone, has been shown in a recent study to be fairly young, only about 8 million years old. The study concludes that the Drake Passage would be free to allow significant deep water flow by around 31 Ma. This would have facilitated an earlier onset of the Antarctic Circumpolar Current. Currently, an opening of the Drake Passage during the early Oligocene is favored.


The opening of the Tasman Gateway

The other major oceanic gateway opening during this time was the Tasman, or Tasmanian, depending on the paper, gateway between Australia and Antarctica. The time frame for this opening is less disputed than the Drake Passage and is largely considered to have occurred around 34 Ma. As the gateway widened, the Antarctic Circumpolar Current strengthened.


The Tethys Seaway closing

The
Tethys Seaway The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
was not a gateway, but rather a sea in its own right. Its closing during the Oligocene had significant impact on both ocean circulation and climate. The collisions of the African plate with the European plate and of the Indian subcontinent with the Asian plate, cut off the Tethys Seaway that had provided a low-latitude ocean circulation. The closure of Tethys built some new mountains (the Zagros range) and drew down more carbon dioxide from the atmosphere, contributing to global cooling.


Greenland–Iceland–Faroes

The gradual separation of the clump of continental crust and the deepening of the tectonic ridge in the North Atlantic that would become Greenland, Iceland, and the Faroe Islands helped to increase the deep water flow in that area. More information about the evolution of North Atlantic Deep Water will be given a few sections down.


Ocean cooling

Evidence for ocean-wide cooling during the Oligocene exists mostly in isotopic proxies. Patterns of extinction and patterns of species migration can also be studied to gain insight into ocean conditions. For a while, it was thought that the glaciation of Antarctica may have significantly contributed to the cooling of the ocean, however, recent evidence tends to deny this.


Deep water

Isotopic evidence suggests that during the early Oligocene, the main source of deep water was the North Pacific and the
Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-smal ...
. As the Greenland-Iceland-Faroe Ridge sank and thereby connected the Norwegian–Greenland sea with the Atlantic Ocean, the deep water of the
North Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe a ...
began to come into play as well. Computer models suggest that once this occurred, a more modern in appearance thermo-haline circulation started.


North Atlantic deep water

Evidence for the early Oligocene onset of chilled North Atlantic deep water lies in the beginnings of sediment drift deposition in the North Atlantic, such as the Feni and Southeast Faroe drifts.


South Ocean deep water

The chilling of the South Ocean deep water began in earnest once the Tasmanian Gateway and the Drake Passage opened fully. Regardless of the time at which the opening of the Drake Passage occurred, the effect on the cooling of the Southern Ocean would have been the same.


Impact events

Recorded extraterrestrial impacts: * Haughton impact crater,
Nunavut Nunavut ( , ; iu, ᓄᓇᕗᑦ , ; ) is the largest and northernmost territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the '' Nunavut Act'' and the '' Nunavut Land Claims Agreement Act'' ...
, Canada (23 Ma, crater diameter) (now considered questionable as an Oligocene event; later analyses have concluded the crater dates to 39 Ma, placing the event in the Eocene.)


Supervolcanic explosions

* La Garita Caldera (28–26 million years ago) * Wah Wah Springs Caldera (30 million years ago)


See also

* List of fossil sites ''(with link directory)'' *
Turgai Sea The Turgai Sea, also known as the Turgay Sea, Turgai Strait, Obik Sea, Ural Sea or West Siberian Sea, was a large shallow body of salt water (an epicontinental or epeiric sea) during the Mesozoic through Cenozoic Eras. It extended north of the pre ...


References

* Ogg, Jim; June, 2004, ''Overview of Global Boundary Stratotype Sections and Points (GSSP's)'

Accessed April 30, 2006.


External links


Palaeos: OligoceneOligocene Microfossils: 300+ images of Foraminifera
{{Authority control Oligocene, Geological epochs Paleogene geochronology