Olduvai Hominid 9
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Olduvai Hominid number 9 (OH 9) is a fossilized skull cap of an early
hominin The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The ...
, found in LLK II,
Olduvai Gorge The Olduvai Gorge or Oldupai Gorge in Tanzania is one of the most important paleoanthropological localities in the world; the many sites exposed by the gorge have proven invaluable in furthering understanding of early human evolution. A steep-si ...
by Louis S. B. Leakey in 1960. It is believed to be ca. 1.4 million years old. Its cranial capacity is estimated at than , the largest value among all known African '' Homo erectus'' specimens. OH 9 is significant because the features it carried and its correlation to the species classification it resides in.


Species Classification

Leakey named it "Chellean Man", in reference to the
Oldowan The Oldowan (or Mode I) was a widespread stone tool archaeological industry (style) in prehistory. These early tools were simple, usually made with one or a few flakes chipped off with another stone. Oldowan tools were used during the Lower ...
tools found at the site, which were then referred to by the now-obsolete name Chellean. Heberer (1963) provisionally named a new species ''Homo leakeyi'' based on the specimen in honor of Leakey, but most subsequent workers have regarded it as ''
Homo ergaster ''Homo ergaster'' is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether ''H. ergaster'' constitutes a species of its own or should be subsumed into '' H. erectus'' is an ongoing and unresol ...
'', or as '' Homo erectus'' (''H. ergaster'' is sometimes regarded as a subspecies of ''H. erectus'', ''H. erectus ergaster'').
Phillip Tobias Phillip Vallentine Tobias (14 October 1925 – 7 June 2012) was a South African palaeoanthropologist and Professor Emeritus at the University of the Witwatersrand in Johannesburg. He was best known for his work at South Africa's hominid fossil ...
provisionally named a new subspecies, ''H. erectus olduvaiensis'', in 1968 based on the specimen, but this has not seen continued use. To the extent that proponents of the use of ''H. ergaster'' define ''ergaster'' as a separate species (instead of an "African ''H. erectus''") rather than a pure
chronospecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
, the assignment of OH 9 to ''H. erectus sensu stricto'' by
Colin Groves Colin Peter Groves (24 June 1942 – 30 November 2017) was a British-Australian biologist and anthropologist. Groves was Professor of Biological Anthropology at the Australian National University in Canberra, Australia. Education Born in Englan ...
supports subsuming ''H. ergaster'' into ''H. erectus''. Because the name ''Homo leakeyi'' was proposed conditionally, it is not valid according to the rules established by the ICZN. Kretzoi (1984) created the replacement name ''Homo louisleakeyi'', which is valid. Cranial bone thickness has been widely qualified for ''Homo erectus'' but rarely quantified. It’s quite often that throughout craniums found, the thickness varies between those different hominids. Yet in OH 9, compared to other ''H. erectus'', it had the biggest cranial capacity standing at 1,067 cc and one of the largest mid supra-orbital torus thickness of 18.5 mm also known as the brow ridge. OH 9 has a robust brow ridge that allows it to stand out among other ''H. erectus''. The brow ridge made it difficult to determine whether this cranium should be classified as ''H. erectus'' or a different species. Cranial bone thickness is key when determining whether a specimen found is ''H. erectus''. The pattern of bone thickness distribution observed in Asian ''H. erectus'', ''P. paniscus'', and possibly in the australopiths, early ''Homo'' or African ''H. ergaster/erectus'' analyzed appears to be a pleomorphic trait among hominids. Since cranium thickness and the OH 9 cranium capacity is larger than any found, it explored the idea of different forms of ''H. erectus'' from different areas due to migration. By the 1980s, the growing numbers of ''H. erectus'' specimens, particularly in Africa, led to the realization that Asian ''Homo erectus'' (''H. erectus sensu stricto''), once thought so primitive, was in fact more derived than its African counter-parts, which leads into the
Out of Africa hypothesis In paleoanthropology, the recent African origin of modern humans, also called the "Out of Africa" theory (OOA), recent single-origin hypothesis (RSOH), replacement hypothesis, or recent African origin model (RAO), is the dominant model of the ...
of humans origins.


Cranial Features

The significant features of the OH 9 also consisted of the structure of the skull. O.H. 9 had a flatter frontal squama curving not as steep from a broader supratoral shelf. Also, OH 9 showed no sign of keeling in the midline, which was often found in ''H. erectus'' especially with a protruding brow ridge. The occipital torus thickness also known as the occipital bun which is the back of the head was also fairly thick compared to most ''H. erectus'' with a thickness of 18.5 mm. OH 9 being the largest cranium capacity, helped us learn the increase of brain growth in ''H. erectus''. Which implied that major differences in the development of cognitive capabilities existed between ''H. erectus'' and anatomically modern humans. These new traits like the robust brow ridge and skull thickness confirmed that ''H. erectus'' in different areas kept evolving. In the same site, the oldowan tools found could be assumed to have been used by this exact ''H. erectus''. This allowed anthropologist to understand how advanced this unique ''H. erectus'' cognitive behavior truly was.


References


External links

*{{commons category-inline, OH 9 Homo erectus fossils Fossils of Tanzania