Oceanian realm
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The Oceanian realm is one of the
World Wildlife Fund The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wo ...
(WWF)
biogeographic realm A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions. De ...
s, and is unique in not including any
continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions are commonly regarded as continents. Ordered from largest in area to smallest, these seven ...
al land mass. It has the smallest land area of any of the WWF realms. This realm includes the islands of the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
in
Micronesia Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, and ...
, the Fijian Islands, the
Hawaiian islands The Hawaiian Islands ( haw, Nā Mokupuni o Hawai‘i) are an archipelago of eight major islands, several atolls, and numerous smaller islets in the North Pacific Ocean, extending some from the island of Hawaii in the south to northernmost Kur ...
, and
Polynesia Polynesia () "many" and νῆσος () "island"), to, Polinisia; mi, Porinihia; haw, Polenekia; fj, Polinisia; sm, Polenisia; rar, Porinetia; ty, Pōrīnetia; tvl, Polenisia; tkl, Polenihia (, ) is a subregion of Oceania, made up of ...
(with the exception of New Zealand).
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, and most of
Melanesia Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from Indonesia's New Guinea in the west to Fiji in the east, and includes the Arafura Sea. The region includes the four independent countries of Fiji, Va ...
including
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
,
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...
, the
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...
, and
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
are included within the
Australasian realm The Australasian realm is a biogeographic realm that is coincident with, but not (by some definitions) the same as, the geographical region of Australasia. The realm includes Australia, the island of New Guinea (comprising Papua New Guinea and ...
. Conversely, New Guinea, New Caledonia, the Solomon Islands and New Zealand are included in the Oceanian realm in the
classification scheme In information science and ontology, a classification scheme is the product of arranging things into kinds of things (classes) or into ''groups'' of classes; this bears similarity to categorization, but with perhaps a more theoretical bent, as cla ...
developed by
Miklos Udvardy Miklos Dezso Ferenc Udvardy (March 23, 1919 – January 27, 1998) was a Hungarian-American biologist and university instructor. He contributed to a wide variety of disciplines, such as biogeography, evolutionary biology, ornithology, and veget ...
in 1975.Udvardy, M. D. F. (1975). ''A classification of the biogeographical provinces of the world''. IUCN Occasional Paper no. 18. Morges, Switzerland: IUCN.Udvardy, Miklos D. F. (1975) ''World Biogeographical Provinces'' (Map). The CoEvolution Quarterly, Sausalito, California
link
The
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
have been included in both the Oceanian and
Temperate South America Temperate South America is a biogeography, biogeographic region of the Earth's seas, comprising the temperate and subtropical waters of South America, including both the Pacific Ocean, Pacific and Atlantic Ocean, Atlantic coasts of the continent a ...
n realms. Despite only being a few hundred miles removed from the South American coast, the islands have strong Hawaiian and southeast
Polynesia Polynesia () "many" and νῆσος () "island"), to, Polinisia; mi, Porinihia; haw, Polenekia; fj, Polinisia; sm, Polenisia; rar, Porinetia; ty, Pōrīnetia; tvl, Polenisia; tkl, Polenihia (, ) is a subregion of Oceania, made up of ...
n biogeographic influences, and the presence of an endemic insect and plant family. The uninhabited French territory of
Clipperton Island Clipperton Island ( or ; ) is an uninhabited, coral atoll in the eastern Pacific Ocean. It is from Paris, France, from Papeete, Tahiti, and from Mexico. It is an Overseas France, overseas state private property of France under direct authori ...
, 1,000 kilometers off the coast of
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, has also been associated with the realm.


Geology

Oceania is geologically the youngest realm. While other realms include old continental land masses or fragments of continents, Oceania is composed mostly of volcanic
high islands Geologically, a high island or volcanic island is an island of volcanic origin. The term can be used to distinguish such islands from low islands, which are formed from sedimentation or the uplifting of coral reefs (which have often formed ...
and
coral atoll Corals are marine invertebrates within the class (biology), class Anthozoa of the phylum Cnidaria. They typically form compact Colony (biology), colonies of many identical individual polyp (zoology), polyps. Coral species include the important C ...
s that arose from the sea in geologically recent times, many of them in the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
. They were created either by hotspot volcanism, or as
island arc Island arcs are long chains of active volcanoes with intense seismic activity found along convergent tectonic plate boundaries. Most island arcs originate on oceanic crust and have resulted from the descent of the lithosphere into the mantle alon ...
s pushed upward by the collision and
subduction Subduction is a geological process in which the oceanic lithosphere is recycled into the Earth's mantle at convergent boundaries. Where the oceanic lithosphere of a tectonic plate converges with the less dense lithosphere of a second plate, the ...
of
tectonic plate Plate tectonics (from the la, label=Late Latin, tectonicus, from the grc, τεκτονικός, lit=pertaining to building) is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large te ...
s. The islands range from tiny islets, sea stacks and coral
atoll An atoll () is a ring-shaped island, including a coral rim that encircles a lagoon partially or completely. There may be coral islands or cays on the rim. Atolls are located in warm tropical or subtropical oceans and seas where corals can gr ...
s to large mountainous islands, like
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
and
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
.


Climate

The climate of Oceania's islands is tropical or subtropical, and range from humid to seasonally dry. Wetter parts of the islands are covered by
tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discont ...
, while the drier parts of the islands, including the leeward sides of the islands and many of the low coral islands, are covered by
tropical and subtropical dry broadleaf forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...
and
Tropical and subtropical grasslands, savannas, and shrublands Tropical and subtropical grasslands, savannas, and shrublands is a terrestrial biome defined by the World Wide Fund for Nature. The biome is dominated by grass and/or shrubs located in semi-arid to semi-humid climate regions of subtropical and t ...
.
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
's high volcanoes,
Mauna Kea Mauna Kea ( or ; ; abbreviation for ''Mauna a Wākea''); is a dormant volcano on the island of Hawaii. Its peak is above sea level, making it the highest point in the state of Hawaii and second-highest peak of an island on Earth. The peak is ...
and
Mauna Loa Mauna Loa ( or ; Hawaiian: ; en, Long Mountain) is one of five volcanoes that form the Island of Hawaii in the U.S. state of Hawaii in the Pacific Ocean. The largest subaerial volcano (as opposed to subaqueous volcanoes) in both mass and ...
, are home to some rare tropical
montane grasslands and shrublands Montane grasslands and shrublands is a biome defined by the World Wildlife Fund. The biome includes high elevation grasslands and shrublands around the world. The term "montane" in the name of the biome refers to "high elevation", rather than th ...
.


Flora and fauna

Since the islands of Oceania were never connected by land to a continent, the flora and fauna of the islands originally reached them from across the ocean (though at the height of the last ice age sea levels were much lower than today and many current
seamount A seamount is a large geologic landform that rises from the ocean floor that does not reach to the water's surface (sea level), and thus is not an island, islet or cliff-rock. Seamounts are typically formed from extinct volcanoes that rise abru ...
s were islands, so some now isolated islands were once less isolated). Once they reached the islands, the ancestors of Oceania's present flora and fauna adapted to life on the islands. Larger islands with diverse ecological niches encouraged floral and faunal
adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
, whereby multiple species evolved from a common ancestor, each species adapted to a different ecological niche; the various species of
Hawaiian honeycreeper Hawaiian honeycreepers are a group of small, passerine birds endemic to Hawaii. They are closely related to the rosefinches in the genus ''Carpodacus'', but many species have evolved features unlike those present in any other finch. Their great ...
s (Family ''Drepanididae'') are a classic example. Other adaptations to island ecologies include gigantism, dwarfism, and among birds, loss of flight. Oceania has a number of
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
species; Hawaii in particular is considered a global
center of endemism A Centre of Endemism is an area in which the ranges of restricted-range species overlap, or a localised area which has a high occurrence of endemics. Centres of endemism may overlap with biodiversity hotspots which are biogeographic regions charact ...
, with its forest
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...
s having one of the highest percentages of endemic plants in the world.


Flora

Land
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s disperse by several different means. Many plants, mostly ferns and mosses but also some flowering plants, disperse on the wind, relying on tiny spores or feathery seeds that can remain airborne over long distances notably ''
Metrosideros ''Metrosideros'' is a genus of approximately 60 trees, shrubs, and vines mostly found in the Pacific region in the family Myrtaceae. Most of the tree forms are small, but some are exceptionally large, the New Zealand species in particular. The n ...
'' trees from New Zealand spread on the wind across Oceania. Other plants, notably
coconut palm The coconut tree (''Cocos nucifera'') is a member of the palm tree family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the f ...
s and
mangroves A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in several ...
, produce seeds that can float in salt water over long distances, eventually washing up on distant beaches, and thus '' Cocos'' trees are ubiquitous across Oceania.
Bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s are also an important means of dispersal; some plants produce sticky seeds that are carried on the feet or feathers of birds, and many plants produce fruits with seeds that can pass through the digestive tracts of birds. ''
Pandanus ''Pandanus'' is a genus of monocots with some 750 accepted species. They are palm-like, dioecious trees and shrubs native to the Old World tropics and subtropics. The greatest number of species are found in Madagascar and Malaysia. Common names ...
'' trees are fairly ubiquitous across Oceania. Botanists generally agree that much of the flora of Oceania is derived from the
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. The ...
n Flora of the
Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
, and
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
, with some plants from Australasia and a few from the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
, particularly in Hawaii.
Easter Island Easter Island ( rap, Rapa Nui; es, Isla de Pascua) is an island and special territory of Chile in the southeastern Pacific Ocean, at the southeasternmost point of the Polynesian Triangle in Oceania. The island is most famous for its nearl ...
has some plants from
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
such as the totora reed.


Fauna

Dispersal across the ocean is difficult for most land
animal Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motilit ...
s, and Oceania has relatively few indigenous land animals compared to other realms. Certain types of animals that are ecologically important on the continental realms, like large land
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s and grazing mammals, were entirely absent from the islands of Oceania until humans brought them.
Bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s are relatively common, including many
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s and some species of land birds whose ancestors may have been blown out to sea by storms. Some birds evolved into flightless species after their ancestors arrived, including several species of
rails Rail or rails may refer to: Rail transport *Rail transport and related matters *Rail (rail transport) or railway lines, the running surface of a railway Arts and media Film * ''Rails'' (film), a 1929 Italian film by Mario Camerini * ''Rail'' ( ...
. A number of islands have indigenous
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s, including
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from . Geckos ar ...
es and
skink Skinks are lizards belonging to the family Scincidae, a family in the infraorder Scincomorpha. With more than 1,500 described species across 100 different taxonomic genera, the family Scincidae is one of the most diverse families of lizards. Ski ...
s, whose ancestors probably arrived on floating rafts of vegetation washed out to sea by storms. With the exception of
bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most ...
s, which live on most of the island groups, there are few if any indigenous non-marine
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s in Oceania.


Impact of settlement

Many animal and plant species have been introduced by humans in two main waves.
Malayo-Polynesian The Malayo-Polynesian languages are a subgroup of the Austronesian languages, with approximately 385.5 million speakers. The Malayo-Polynesian languages are spoken by the Austronesian peoples outside of Taiwan, in the island nations of Southeast ...
settlers brought
pig The pig (''Sus domesticus''), often called swine, hog, or domestic pig when distinguishing from other members of the genus '' Sus'', is an omnivorous, domesticated, even-toed, hoofed mammal. It is variously considered a subspecies of ''Sus ...
s,
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Do ...
s,
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adult m ...
s and
polynesian rat The Polynesian rat, Pacific rat or little rat (''Rattus exulans''), known to the Māori as ''kiore'', is the third most widespread species of rat in the world behind the brown rat and black rat. The Polynesian rat originated in Southeast Asia, a ...
s to many islands; and had spread across the whole of Oceania by 1200 CE. From the seventeenth century onwards European settlers brought other animals, including
cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
s,
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
,
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
s,
small Asian mongoose Small Asian mongoose is a common name applied to two mammals which were formerly considered to be a single species: * Javan mongoose *Small Indian mongoose The small Indian mongoose (''Urva auropunctata'') is a mongoose species native to Iraq an ...
(''Herpestes javanicus''),
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
,
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s, and the
brown rat The brown rat (''Rattus norvegicus''), also known as the common rat, street rat, sewer rat, wharf rat, Hanover rat, Norway rat, Norwegian rat and Parisian rat, is a widespread species of common rat. One of the largest muroids, it is a brown o ...
(''Rattus norvegicus''). These and other introduced species, in addition to overhunting and deforestation, have dramatically altered the ecology of many of Oceania's islands, pushing many species to extinction or near-extinction, or confining them to small islets uninhabited by humans. The absence of predator species caused many bird species to lose the instinct to flee from predators and to lay their eggs on the ground, which makes them vulnerable to predators such as cats, dogs, mongooses or rats. The arrival of humans on these island groups often resulted in disruption of the indigenous ecosystems and waves of species extinctions (see
Holocene extinction event The Holocene extinction, or Anthropocene extinction, is the ongoing extinction event during the Holocene epoch. The extinctions span numerous families of bacteria, fungi, plants, and animals, including mammals, birds, reptiles, amphibians, fish, ...
).
Easter Island Easter Island ( rap, Rapa Nui; es, Isla de Pascua) is an island and special territory of Chile in the southeastern Pacific Ocean, at the southeasternmost point of the Polynesian Triangle in Oceania. The island is most famous for its nearl ...
, the easternmost island in Polynesia, shows evidence of
ecosystem collapse An ecosystem is considered collapsed when its unique biotic (characteristic biota) or abiotic features are lost from all previous occurrences. Ecosystem collapse causes ecological collapse within a system; essentially altering its stability, re ...
, probably caused by human activity, several hundred years ago, which contributed (along with slave raiding and European diseases) to a 99% decline in the human population of the island. The island, once lushly forested, is now mostly windswept grassland. More recently,
Guam Guam (; ch, Guåhan ) is an organized, unincorporated territory of the United States in the Micronesia subregion of the western Pacific Ocean. It is the westernmost point and territory of the United States (reckoned from the geographic cent ...
's native bird and lizard species were decimated after the introduction of the
brown tree snake The brown tree snake (''Boiga irregularis''), also known as the brown catsnake, is an arboreal rear-fanged colubrid snake native to eastern and northern coastal Australia, eastern Indonesia (Sulawesi to Papua), Papua New Guinea, and many islands ...
(''Boiga irregularis'') in the 1940s.


Ecoregions


References

{{DEFAULTSORT:Oceania Realm Biogeographic realms Environment of Oceania * Natural history of Oceania Phytogeography