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The oystercatchers are a group of waders forming the family Haematopodidae, which has a single genus, ''Haematopus''. They are found on coasts worldwide apart from the polar regions and some tropical regions of Africa and South East Asia. The exceptions to this are the Eurasian oystercatcher, the South Island oystercatcher, and the Magellanic oystercatcher, which also breed inland, far inland in some cases. In the past there has been a great deal of confusion as to the species limits, with discrete populations of all black oystercatchers being afforded specific status but pied oystercatchers being considered one single species.Hockey, P (1996). "Family Haematopodidae (Oystercatchers)". In del Hoyo, J.; Elliot, A. & Sargatal, J. (editors). ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
''. Volume 3: ''Hoatzin to Auks''. Lynx Edicions. .


Taxonomy

The genus ''Haematopus'' was introduced in 1758 by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' to accommodate a single species, the Eurasian oystercatcher ''Haematopus ostralegus''. The genus name ''Haematopus'' comes from the Ancient Greek words ''haima'' αἳμα meaning blood, and ''pous'' πούς meaning foot, referring to the red legs of the Eurasian oystercatcher; it had been in use since Pierre Belon in 1555. The family Haematopodidae was introduced (as the subfamily Haematopodinae) by the French naturalist Charles Bonaparte in 1838. The
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
''oystercatcher'' was coined by Mark Catesby in 1731 for the
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
n species '' H. palliatus'', which he described as eating oysters. Yarrell in 1843 established this as the preferred term, replacing the older name ''sea pie''


Description

The different species of oystercatcher show little variation in shape or appearance. They range from in length and in wingspan. The Eurasian oystercatcher is the lightest on average, at , while the sooty oystercatcher is the heaviest, at . The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of all species is either all-black, or black (or dark brown) on top and white underneath. The variable oystercatcher is slightly exceptional in being either all-black or pied. They are large, obvious, and noisy plover-like birds, with massive long orange or red bills used for smashing or prying open
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
s. The bill shape varies between species, according to the diet. Those birds with blade-like bill tips pry open or smash mollusc shells, and those with pointed bill tips tend to probe for
annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
worms. They show sexual dimorphism, with females being longer-billed and heavier than males.


Feeding

The diet of oystercatchers varies with location. Species occurring inland feed upon earthworms and insect larvae. The diet of coastal oystercatchers is more varied, although dependent upon coast type; on estuaries,
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
s,
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
s and polychaete worms are the most important part of the diet, whereas rocky shore oystercatchers prey upon limpets,
mussel Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and Freshwater bivalve, freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other ...
s, gastropods, and
chiton Chitons () are marine molluscs of varying size in the class Polyplacophora (), formerly known as Amphineura. About 940 extant and 430 fossil species are recognized. They are also sometimes known as gumboots or sea cradles or coat-of-mail s ...
s. Other prey items include echinoderms, fish, and crabs.


Breeding

Nearly all species of oystercatcher are
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
, although there are reports of polygamy in the Eurasian oystercatcher. They are
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
during the breeding season (with a few species defending territories year round). There is strong mate and site fidelity in the species that have been studied, with one record of a pair defending the same site for 20 years. A single nesting attempt is made per breeding season, which is timed over the summer months. The nests of oystercatchers are simple affairs, scrapes in the ground which may be lined, and placed in a spot with good visibility. The eggs of oystercatchers are spotted and cryptic. Between one and four eggs are laid, with three being typical in the Northern Hemisphere and two in the south. Incubation is shared but not proportionally, females tend to take more incubation and males engage in more territory defence. Incubation varies by species, lasting between 24–39 days. Oystercatchers are also known to practice "egg dumping." Like the cuckoo, they sometimes lay their eggs in the nests of other species such as seagulls, abandoning them to be raised by those birds.


Conservation

The
Canary Islands oystercatcher The Canary Islands oystercatcher, Canarian oystercatcher, or Canarian black oystercatcher (''Haematopus meadewaldoi''), Etymology: ''Haematopus'', Latinized Ancient Greek for "bloody-footed", from ''(h)aimato-'' (αίματό-), '"blood-" + ''-p ...
became extinct during the 20th century. The
Chatham oystercatcher The Chatham oystercatcher or Chatham Island oystercatcher (''Haematopus chathamensis'') is a species of oystercatcher. It is a wading bird endemic to the Chatham Islands, New Zealand. This species is rated by the IUCN as endangered, and has a cu ...
is endemic to the Chatham Islands of New Zealand but is listed as endangered by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, while both the African and Eurasian oystercatchers are considered near threatened. There has been conflict with commercial shellfish farmers, but studies have found that the impact of oystercatchers is much smaller than that of shore crabs.


Species

The genus contains twelve species. Several fossil species are known, including ''Haematopus sulcatus'' (Brodkorb, 1955) from the
Barstovian The Barstovian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 16,300,000 to 13,600,000 years BP, a period of . It is usua ...
of Florida and Zanclean of North Carolina, and which is evidently considered a synonym of the extant species ''Haematopus palliatus''.Haematopus palliatus Temminick 1820 (American oystercatcher) at FossilWorks
/ref>


References


External links

*ARKive
images and movies of the oystercatcher ''(Haematopus ostralegus)''Oystercatcher videos
on the Internet Bird Collection
LIVE webcam at the seashore in Namsos, Norway 2013
{{Taxonbar, from=Q27515 . Shorebirds Haematopodidae Taxa named by Carl Linnaeus no:Tjeldfamilien nn:Tjeldfamilien