''Ovophis okinavensis'', commonly known as the , Ryukyu Island pit viper, and the Okinawan pitviper, is a

venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...

pitviper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subf ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

found in the

Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yona ...

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

.Gumprecht A, Tillack F, Orlov NL,

Captain A Captain is a title, an appellative for the commanding officer of a military unit; the supreme leader of a navy ship, merchant ship, aeroplane, spacecraft, or other vessel; or the commander of a port, fire or police department, election precinct, e ...

, Ryabov S. 2004. ''Asian Pitvipers''. GeitjeBooks. Berlin. 1st Edition. 368 pp. . No

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

are currently recognized.

Description

Adults are usually 30 to 80 cm (11¾–31½ inches) long. Body usually pale greenish-brown, or yellowish-olive (sometimes pale brown), with alternating, darker brownish or greenish dorsal blotches, each bordered with yellowish scales. Head large, triangular, distinct from neck, narrow dark postocular stripe. Scalation includes: 23 or 21 rows of

dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...

at midbody; 125–135

ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...

; 36–55 paired subcaudal scales; and 8 (sometimes 7 or 9) supralabial scales. The color pattern consists of a gray ground color overlaid with a series of dark gray of grayish-black crossbands. A ventrolateral pattern of black spots against a gray-white background is also present.

Common names

The name Okinawa habu usually refers to Hon habu or simply habu (''

Protobothrops flavoviridis ''Protobothrops flavoviridis'' is a species of venomous pit viper endemic to the Ryukyu Islands of Japan. No subspecies are currently recognized. Local common names include habu,Gumprecht A, Tillack F, , Captain A, Ryabov S. 2004. ''Asian Pitvipe ...

''). The species is also known as kufah.Mehrtens JM. 1987. ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. .Brown JH. 1973. ''Toxicology and Pharmacology of Venoms from Poisonous Snakes''. Springfield, Illinois: Charles C. Thomas. 184 pp. LCCCN 73–229. . This snake is sometimes referred to as in the

Okinawan language The Okinawan language (, , , ) or Central Okinawan, is a Northern Ryukyuan language spoken primarily in the southern half of the island of Okinawa, as well as in the surrounding islands of Kerama, Kumejima, Tonaki, Aguni and a number of sm ...

. It is also called Mamushi, but only in

Amami language The The name ''Amami-guntō'' was standardized on February 15, 2010. Prior to that, another name, ''Amami shotō'' (奄美諸島), was also used. is an archipelago in the Satsunan Islands, which is part of the Ryukyu Islands, and is southwest of ...

, whereas Mamushi is the common name for the unrelated species ''Gloydius blomhoffii''.

Geographic range

The species is found in the

, including

Okinawa is a prefecture of Japan. Okinawa Prefecture is the southernmost and westernmost prefecture of Japan, has a population of 1,457,162 (as of 2 February 2020) and a geographic area of 2,281 km2 (880 sq mi). Naha is the capital and largest city ...

and the

Amami Islands The The name ''Amami-guntō'' was standardized on February 15, 2010. Prior to that, another name, ''Amami shotō'' (奄美諸島), was also used. is an archipelago in the Satsunan Islands, which is part of the Ryukyu Islands, and is southwest of ...

. The type locality given is "Okinawa".McDiarmid RW, Campbell JA, Touré T. 1999. ''Snake Species of the World: A Taxonomic and Geographic Reference, vol. 1.'' Herpetologists' League. 511 pp. (series). (volume).

Habitat

''Ovophis okinavensis'' occurs in various

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s, including open

woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (se ...

forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...

mountain A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited Summit (topography), summit area, and ...

fields Fields may refer to: Music * Fields (band), an indie rock band formed in 2006 * Fields (progressive rock band), a progressive rock band formed in 1971 * ''Fields'' (album), an LP by Swedish-based indie rock band Junip (2010) * "Fields", a song b ...

, in

farming Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled peopl ...

areas with nearby

stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...

pond A pond is an area filled with water, either natural or artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% emergent vegetation helps in distinguishing their ecology from ...

s, and other

water source Water supply is the provision of water by public utilities, commercial organisations, community endeavors or by individuals, usually via a system of pumps and pipes. Public water supply systems are crucial to properly functioning societies. Thes ...

s. It can also be found in human habitations

Feeding

The hime habu hunts for

rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...

s and other

vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...

s in open areas, especially in

sugar cane Sugarcane or sugar cane is a species of (often hybrid) tall, perennial grass (in the genus '' Saccharum'', tribe Andropogoneae) that is used for sugar production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fibrous stalk ...

fields and, sometimes near human habitations.

Reproduction

Both oviparous and

ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...

. Depending on environmental condition, females will either deposit their eggs, or retain them to incubate internally and give birth later to live young.

Venom

The

venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a st ...

of ''Ovophis okinavensis'', like that of most

vipers The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs th ...

, is mainly

hemotoxin Hemotoxins, haemotoxins or hematotoxins are toxins that destroy red blood cells, disrupt blood clotting, and/or cause organ degeneration and generalized tissue damage. The term ''hemotoxin'' is to some degree a misnomer since toxins that damage t ...

with cytotoxicity factors. People are bitten when they step on this sluggish snake at night, or when tending crops by day. Although venom from this snake is not life-threatening usually, people still should seek medical attention promptly if they are bitten. Because of its relatively weak venom, antivenom is not produced.