The osculum (plural "oscula") is an excretory structure in the living

sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...

, a large opening to the outside through which the current of water exits after passing through the

spongocoel A spongocoel (), also called paragaster (or paragastric cavity), is the large, central cavity of sponges. Water enters the spongocoel through hundreds of tiny pores ( ostia) and exits through the larger opening (osculum). Depending on the body pla ...

. Wastes diffuse into the water and the water is pumped through the osculum carrying away with it the sponge's wastes. Sponges pump large volumes of water: typically a volume of water equal to the sponge's body size is pumped every five seconds. The size of the osculum is regulated by contractile

myocyte A muscle cell is also known as a myocyte when referring to either a cardiac muscle cell (cardiomyocyte), or a smooth muscle cell as these are both small cells. A skeletal muscle cell is long and threadlike with many nuclei and is called a muscl ...

s. Its size, in turn, is one of the factors which determines the amount of water flowing through the

. It can be closed completely in response to excess silt in the water.

References

Sponge anatomy {{animal-anatomy-stub