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The European rabbit (''Oryctolagus cuniculus'') or coney is a species of
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit speci ...
native to the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
(including
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
,
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
, and southwestern France), western
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, and the northern
Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through Moroc ...
in northwest Africa. It has been widely introduced elsewhere, often with devastating effects on local
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...
. Its decline in its native range due to
myxomatosis Myxomatosis is a disease caused by ''Myxoma virus'', a poxvirus in the genus ''Leporipoxvirus''. The natural hosts are tapeti (''Sylvilagus brasiliensis'') in South and Central America, and brush rabbits (''Sylvilagus bachmani'') in North Ame ...
,
rabbit calicivirus Rabbit hemorrhagic disease (RHD), also known as viral hemorrhagic disease (VHD), is a highly infectious and lethal form of viral hepatitis that affects European rabbits. Some viral strains also affect hares and cottontail rabbits. Mortality rate ...
, overhunting and habitat loss has caused the decline of the
Iberian lynx The Iberian lynx (''Lynx pardinus'') is a wild cat species endemic to the Iberian Peninsula in southwestern Europe. It is listed as Endangered on the IUCN Red List. In the 20th century, the Iberian lynx population had declined because of overhun ...
(''Lynx pardinus'') and
Spanish imperial eagle The Spanish imperial eagle (''Aquila adalberti''), also known as the Spanish eagle or Adalbert's eagle, is a species of eagle native to the Iberian Peninsula. The binomial commemorates Prince Adalbert of Bavaria. Due to its distinct “epaulet ...
(''Aquila adalberti''). It is known as an
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
because it has been introduced to countries on all continents with the exception of Antarctica, and has caused many problems within the environment and ecosystems; in particular, European rabbits in Australia have had a devastating impact, due in part to the lack of natural predators there. The European rabbit is well known for digging networks of
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
s, called warrens, where it spends most of its time when not feeding. Unlike the related
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The ge ...
s (''Lepus'' spp.), rabbits are
altricial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
, the young being born blind and furless, in a fur-lined nest in the warren, and they are totally dependent upon their mothers. Much of the modern research into wild rabbit behaviour was carried out in the 1960s by two research centres. One was the naturalist
Ronald Lockley Ronald Mathias Lockley (8 November 1903 – 12 April 2000) was a Welsh ornithologist and naturalist. He wrote over fifty books on natural history, including a major study of shearwaters, and many articles. He is perhaps best known for his book ...
, who maintained a number of large enclosures for wild rabbit colonies, with observation facilities, in
Orielton, Pembrokeshire Orielton is a historic country house near Hundleton in Pembrokeshire, Wales. It has been used as a field studies centre for environmental sciences but was put on sale in early 2022 History The first known house at Orielton was a fortified manor b ...
. Apart from publishing a number of scientific papers, he popularised his findings in a book ''The Private Life of the Rabbit'', which is credited by
Richard Adams Richard George Adams (9 May 1920 – 24 December 2016) was an English novelist and writer of the books ''Watership Down'', ''Maia'', ''Shardik'' and ''The Plague Dogs''. He studied modern history at university before serving in the British Army ...
as having played a key role in his gaining "a knowledge of rabbits and their ways" that informed his novel ''
Watership Down ''Watership Down'' is an adventure novel by English author Richard Adams, published by Rex Collings Ltd of London in 1972. Set in Berkshire in southern England, the story features a small group of rabbits. Although they live in their natural ...
''. The other group was the
Commonwealth Scientific and Industrial Research Organisation The Commonwealth Scientific and Industrial Research Organisation (CSIRO) is an Australian Government agency responsible for scientific research. CSIRO works with leading organisations around the world. From its headquarters in Canberra, CSIRO ...
(CSIRO) in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, where numerous studies of the social behavior of wild rabbits were performed. Since the onset of myxomatosis, and the decline of the significance of the rabbit as an agricultural pest, few large-scale studies have been performed and many aspects of rabbit behaviour are still poorly understood.


Naming and etymology

Because of its non-British origin, the species does not have native names in
English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ide ...
or
Celtic Celtic, Celtics or Keltic may refer to: Language and ethnicity *pertaining to Celts, a collection of Indo-European peoples in Europe and Anatolia **Celts (modern) *Celtic languages **Proto-Celtic language * Celtic music *Celtic nations Sports Fo ...
, with the usual terms "cony" and "rabbit" being foreign
loanword A loanword (also loan word or loan-word) is a word at least partly assimilated from one language (the donor language) into another language. This is in contrast to cognates, which are words in two or more languages that are similar because th ...
s. "Rabbit" is also pronounced as ''rabbidge'', ''rabbert'' (
North Devon North Devon is a local government district in Devon, England. North Devon Council is based in Barnstaple. Other towns and villages in the North Devon District include Braunton, Fremington, Ilfracombe, Instow, South Molton, Lynton and Lynmouth ...
) and ''rappit'' (
Cheshire Cheshire ( ) is a ceremonial and historic county in North West England, bordered by Wales to the west, Merseyside and Greater Manchester to the north, Derbyshire to the east, and Staffordshire and Shropshire to the south. Cheshire's county t ...
and
Lancashire Lancashire ( , ; abbreviated Lancs) is the name of a historic county, ceremonial county, and non-metropolitan county in North West England. The boundaries of these three areas differ significantly. The non-metropolitan county of Lancashi ...
). More archaic pronunciations include ''rabbette'' (15th-16th centuries), ''rabet'' (15th-17th centuries), ''rabbet'' (16th-18th centuries), ''rabatte'' (16th century), ''rabytt'' (17th century) and ''rabit'' (18th century). The
root word A root (or root word) is the core of a word that is irreducible into more meaningful elements. In morphology, a root is a morphologically simple unit which can be left bare or to which a prefix or a suffix can attach. The root word is the prima ...
is the Walloon ''rabett'', which was once commonly used in
Liège Liège ( , , ; wa, Lîdje ; nl, Luik ; german: Lüttich ) is a major city and municipality of Wallonia and the capital of the Belgian province of Liège. The city is situated in the valley of the Meuse, in the east of Belgium, not far from b ...
. ''Rabett'' itself is derived from the
Middle Dutch Middle Dutch is a collective name for a number of closely related West Germanic dialects whose ancestor was Old Dutch. It was spoken and written between 1150 and 1500. Until the advent of Modern Dutch after 1500 or c. 1550, there was no overarch ...
''robbe'', with the addition of the
suffix In linguistics, a suffix is an affix which is placed after the stem of a word. Common examples are case endings, which indicate the grammatical case of nouns, adjectives, and verb endings, which form the conjugation of verbs. Suffixes can carry ...
''-ett''. The term "cony" or "coney" antedates "rabbit", and first occurred during the 13th century to refer to the animal's pelt. Later, "cony" referred to the adult animal, while "rabbit" referred to the young. The root of "cony" is the
old French Old French (, , ; Modern French: ) was the language spoken in most of the northern half of France from approximately the 8th to the 14th centuries. Rather than a unified language, Old French was a linkage of Romance dialects, mutually intelligib ...
''connil'' or ''counil'', of which the
Norman Norman or Normans may refer to: Ethnic and cultural identity * The Normans, a people partly descended from Norse Vikings who settled in the territory of Normandy in France in the 10th and 11th centuries ** People or things connected with the Norm ...
plural was ''coniz'', and later ''conis''. Its forerunner is the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
κύνικλος, from which the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''cuniculus'' is derived. The origin of κύνικλος itself is unclear: Ælian, who lived during the third century, linked the word to Celtiberian and later authors relate it to its
Basque Basque may refer to: * Basques, an ethnic group of Spain and France * Basque language, their language Places * Basque Country (greater region), the homeland of the Basque people with parts in both Spain and France * Basque Country (autonomous co ...
name ''unchi''; Varo and
Pliny Pliny may refer to: People * Pliny the Elder (23–79 CE), ancient Roman nobleman, scientist, historian, and author of ''Naturalis Historia'' (''Pliny's Natural History'') * Pliny the Younger (died 113), ancient Roman statesman, orator, w ...
connected it to ''cuneus'', which refers to a wedge, thus making reference to the animal's digging ability. The species' dwelling place is termed a warren or cony-garth. "Warren" comes from the Old English ''wareine'', itself derived from the Old French ''warenne'', ''varenne'', or ''garenne''. The root word is the
Low Latin Late Latin ( la, Latinitas serior) is the scholarly name for the form of Literary Latin of late antiquity.Roberts (1996), p. 537. English dictionary definitions of Late Latin date this period from the , and continuing into the 7th century in ...
''warenna'', which originally signified a preserve in general, only to be later used to refer specifically to an enclosure set apart for rabbits and hares. "Cony-garth" derives from the
Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English p ...
''conygerthe'', which may be a
compound Compound may refer to: Architecture and built environments * Compound (enclosure), a cluster of buildings having a shared purpose, usually inside a fence or wall ** Compound (fortification), a version of the above fortified with defensive struct ...
of ''connynge''+''erthe'' (cony+earth). The term stems from the Old French ''conniniere'' or ''coninyere'', and later ''conilliere''. The root word is the Low Latin ''cunicularia'', the feminine form of the
adjective In linguistics, an adjective (list of glossing abbreviations, abbreviated ) is a word that generally grammatical modifier, modifies a noun or noun phrase or describes its referent. Its semantic role is to change information given by the noun. Tra ...
''cunicularius'', which pertains to the rabbit.


Taxonomy

Originally assigned to the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Lepus Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The gen ...
'', the European rabbit was consigned to its own genus in 1874 on account of its
altricial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
young, its burrowing habits, and numerous skeletal characters. It is superficially similar to the North American
cottontail Cottontail rabbits are the leporid species in the genus ''Sylvilagus'', found in the Americas. Most ''Sylvilagus'' species have stub tails with white undersides that show when they retreat, giving them their characteristic name. However, this ...
s (''Sylvilagus'') in that they are born blind and naked, have white flesh, and little
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
. However, they differ in skull characteristics, and cottontails do not habitually construct their own burrows as the European rabbit does. Molecular studies confirm that the resemblance between the two is due to
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
, and that the European rabbit's closest relatives are the
hispid hare The hispid hare (''Caprolagus hispidus''), also called Assam rabbit and bristly rabbit, is a leporid native to South Asia, whose historic range extended along the southern foothills of the Himalayas. Today, its habitat is highly fragmented with an ...
, the riverine rabbit, and the
Amami rabbit The Amami rabbit (''Pentalagus furnessi''; ), or , also known as the Ryukyu rabbit is a dark-furred rabbit which is only found in Amami Ōshima and Toku-no-Shima, two small islands between southern Kyūshū and Okinawa in Kagoshima Prefecture ...
. The oldest known fossils attributed to the modern European rabbit species are around 0.5 Ma old (Middle
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
). The
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
is from Matthee et al., 2004, based on nuclear and mitochondrial gene analysis.


Subspecies

, six subspecies are recognised by MSW3. Genetic studies undertaken in 2008, however, indicate only two subspecies, ''O. c. algirus'' and ''O. c. cuniculus'', with a hybrid zone connecting the two populations in central Iberia.


Description

The European rabbit is smaller than the
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
and
mountain hare The mountain hare (''Lepus timidus''), also known as blue hare, tundra hare, variable hare, white hare, snow hare, alpine hare, and Irish hare, is a Palearctic hare that is largely adapted to polar and mountainous habitats. Evolution The mountai ...
, and lacks black ear tips, as well as having proportionately shorter legs. An adult European rabbit can measure in length, and weigh . The hind foot measures in length, while the ears are long from the
occiput The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone overlies the occipital lobes of the cereb ...
. Size and weight vary according to food and habitat quality, with rabbits living on light soil with nothing but grass to feed on being noticeably smaller than specimens living on highly cultivated farmlands with plenty of roots and clover. Pure European rabbits weighing and upwards are uncommon, but are occasionally reported. One large specimen, caught in February 1890 in
Lichfield Lichfield () is a cathedral city and civil parish in Staffordshire, England. Lichfield is situated roughly south-east of the county town of Stafford, south-east of Rugeley, north-east of Walsall, north-west of Tamworth and south-west of B ...
, was weighed at . Unlike the brown hare, the male European rabbit is more heavily built than the female. The
penis A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males do n ...
is short, and lacks a
baculum The baculum (also penis bone, penile bone, or ''os penis'', ''os genitale'' or ''os priapi'') is a bone found in the penis of many placental mammals. It is absent from the human penis, but present in the penises of some primates, such as the ...
and true
glans The glans (, plural "glandes" ; from the Latin word for "acorn") is a vascular structure located at the tip of the penis in male mammals or a homologous genital structure of the clitoris in female mammals. Structure The exterior structure ...
. The fur of the European rabbit is generally greyish-brown, but this is subject to much variation. The
guard hair Guard hair or overhair is the outer layer of hair of most mammals, which overlay the fur. Guard hairs are long and coarse and protect the rest of the pelage (fur) from abrasion and frequently from moisture. They are visible on the surface of the fu ...
s are banded brown and black, or grey, while the
nape The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , "spinal marrow"). The corresponding adjective is ''nuchal'', as in the term ''nu ...
of the neck and
scrotum The scrotum or scrotal sac is an anatomical male reproductive structure located at the base of the penis that consists of a suspended dual-chambered sac of skin and smooth muscle. It is present in most terrestrial male mammals. The scrotum cont ...
are reddish. The chest patch is brown, while the rest of the underparts are white or grey. A white star shape is often present on kits' foreheads, but rarely occurs in adults. The
whiskers Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to touch, sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are c ...
are long and black, and the feet are fully furred and buff-coloured. The tail has a white underside, which becomes prominent when escaping danger. This may act as a signal for other rabbits to run.
Moulting In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
occurs once a year, beginning in March on the face and spreading over the back. The underfur is completely replaced by October–November. The European rabbit exhibits great variation in colour, from light sandy, to dark grey and completely black. Such variation depends largely on the amount of guard hairs relative to regular pelage. Melanists are not uncommon in
mainland Europe Continental Europe or mainland Europe is the contiguous continent of Europe, excluding its surrounding islands. It can also be referred to ambiguously as the European continent, – which can conversely mean the whole of Europe – and, by ...
, though
albino Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and pink or blue eyes. Individuals with the condition are referred to as albino. Varied use and interpretation of the term ...
es are rare.


Life history and behaviour


Social and territorial behaviours

The European rabbit lives in warrens that contain 2-10 other individuals living in smaller groups to ensure greater breeding success.
Territoriality In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...
and aggression contribute greatly to the rabbits' maturation process, and help ensure survival of the population. Females tend to be more territorial than males, although the areas most frequented by females are not defended. Territories are marked with dung hills. The size of the species'
home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. He ...
varies according to habitat, food, shelter, cover from predators, and breeding sites, though it is generally small, encompassing about 0.3-0.7
hectare The hectare (; SI symbol: ha) is a non-SI metric unit of area equal to a square with 100-metre sides (1 hm2), or 10,000 m2, and is primarily used in the measurement of land. There are 100 hectares in one square kilometre. An acre is a ...
s. Except during times of low rabbit density and abundance of high-quality food, male ranges tend to be larger than those held by females. The European rabbit rarely strays far from its burrow; when feeding on cultivated fields, it typically only moves 25 m away from its burrow, and rarely 50 m. It may, however, move as far as 500 m after an abrupt change in environment, such as a
harvest Harvesting is the process of gathering a ripe crop from the fields. Reaping is the cutting of grain or pulse for harvest, typically using a scythe, sickle, or reaper. On smaller farms with minimal mechanization, harvesting is the most labor-i ...
. This behaviour may be an antipredator adaptation, as rabbits in areas where predators are under rigorous control may move three times further from their burrows than those in areas without predator management. The European rabbit is a gregarious animal, which lives in stable social groups centred around females sharing access to one or more burrow systems. Social structures tend to be looser in areas where burrow construction is relatively easy.
Dominance hierarchies In biology, a dominance hierarchy (formerly and colloquially called a pecking order) is a type of social hierarchy that arises when members of animal social groups interact, creating a ranking system. A dominant higher-ranking individual is som ...
exist in parallel for both bucks and does. Among bucks, status is determined through access to does, with dominant bucks siring the majority of the colony's offspring. The dominant does have priority access to the best nesting sites, with competition over such sites often leading to serious injury or death. Subordinate does, particularly in large colonies, typically resort to using single-entrance breeding spots far from the main warren, thus making themselves vulnerable to
fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
or
badger Badgers are short-legged omnivores in the family Mustelidae (which also includes the otters, wolverines, martens, minks, polecats, weasels, and ferrets). Badgers are a polyphyletic rather than a natural taxonomic grouping, being united b ...
predation.


Reproduction and development

In the European rabbit's
mating system A mating system is a way in which a group is structured in relation to sexual behaviour. The precise meaning depends upon the context. With respect to animals, the term describes which males and females mating, mate under which circumstances. Reco ...
, dominant bucks exhibit
polygyny Polygyny (; from Neoclassical Greek πολυγυνία (); ) is the most common and accepted form of polygamy around the world, entailing the marriage of a man with several women. Incidence Polygyny is more widespread in Africa than in any o ...
, whereas lower-status individuals (both bucks and does) often form
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
breeding relationships. Rabbits signal their readiness to copulate by marking other animals and inanimate objects with an odoriferous substance secreted though a chin gland, in a process known as "chinning". Though male European rabbits may sometimes be amicable with one another, fierce fights can erupt among bucks during the
breeding season Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and cha ...
, typically January to August. A succession of litters (usually three to seven kittens each) are produced, but in overpopulated areas, pregnant does may lose all their
embryo An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male spe ...
s through intrauterine resorption. Shortly before giving birth, the doe constructs a separate burrow known as a "stop" or "stab", generally in an open field away from the main warren. These breeding burrows are typically a few feet long and are lined with grass and moss, as well as fur plucked from the doe's belly. The breeding burrow protects the kits from adult bucks and predators. The
gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once it ...
of the European rabbit is 30 days, with the
sex ratio The sex ratio (or gender ratio) is usually defined as the ratio of males to females in a population. As explained by Fisher's principle, for evolutionary reasons this is typically about 1:1 in species which reproduce sexually. Many species devia ...
of male to female kits tending to be 1:1. Greater maternal investment over male offspring may result in higher
birth weight Birth weight is the body weight of a baby at its birth. The average birth weight in babies of European descent is , with the normative range between . On average, babies of South Asian and Chinese descent weigh about . As far as low birth weight ...
s for bucks. Kits born to the dominant buck and doe—which enjoy better nesting and feeding grounds—tend to grow larger and stronger and to become more dominant than those born to subordinate rabbits. Not uncommonly, European rabbits mate again immediately after giving birth, with some specimens having been observed to nurse previous young whilst pregnant. Female European rabbits nurse their kits once a night, for only a few minutes. After suckling is complete, the doe seals the entrance to the stop with soil and vegetation. In its native Iberian and southern French range, European rabbit young have a growth rate of per day, though such kittens in non-native ranges may grow per day. Weight at birth is and increases to by 21–25 days, during the
weaning Weaning is the process of gradually introducing an infant human or another mammal to what will be its adult diet while withdrawing the supply of its mother's milk. The process takes place only in mammals, as only mammals produce milk. The infan ...
period. European rabbit kits are born blind, deaf, and nearly naked. The ears do not gain the power of motion until 10 days of age, and can be erected after 13. The eyes open 11 days after birth. At 18 days, the kittens begin to leave the burrow.
Sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definitio ...
in bucks is attained at 4 months, while does can begin to breed at 3-5 months.


Burrowing behaviour

The European rabbit's burrows occur mostly on slopes and banks, where
drainage Drainage is the natural or artificial removal of a surface's water and sub-surface water from an area with excess of water. The internal drainage of most agricultural soils is good enough to prevent severe waterlogging (anaerobic conditio ...
is more efficient. The burrow entrances are typically 10–50 cm in diameter, and are easily recognisable by the bare earth at their mouths. Vegetation growth is prevented by the constant passing and repassing of the resident rabbits. Big burrows are complex excavations which may descend to depths of several feet. They are not constructed on any specified plan, and appear to be enlarged or improved as a result of the promiscuous activity of several generations. Digging is done by pulling the soil backwards with the fore feet and throwing it between the hind legs, which scatter the material with kicking motions. While most burrows are dug from the outside, some warrens feature holes dug from the inside, which act as emergency exits when escaping from predators below ground. These holes usually descend perpendicularly to 3–4 feet, and their mouths lack the bare-earth characteristic of burrow entrances. While kits sleep in chambers lined with grass and fur, adults sleep on the bare earth, likely to prevent damp, with warmth being secured by
huddling In biology, kleptothermy is any form of thermoregulation by which an animal shares in the metabolic thermogenesis of another animal. It may or may not be reciprocal, and occurs in both endotherms and ectotherms. One of its forms is huddling. Howe ...
. Although both sexes dig, does do so more skillfully, and for longer periods.


Communication

The European rabbit is a relatively quiet animal, though it has at least two vocalisations. The best-known is a high treble scream or squeal. This distress call has been likened to the cry of a piglet. This sound is uttered when in extreme distress, such as being caught by a predator or trap. During the spring, bucks express contentment by emitting grunting sounds when approaching other rabbits. These grunts are similar to shrill
hiccup A hiccup (scientific name ''singultus'', from a Latin word meaning "to catch one's breath while sobbing"; also spelled hiccough) is an involuntary contraction (myoclonic jerk) of the diaphragm that may repeat several times per minute. The hic ...
s, and are emitted with the mouth closed. Aggression is expressed with a low growl.


Ecology


Habitat

The European rabbit's ideal habitat consists of short grasslands with secure refuge (such as burrows, boulders, hedgerows, scrub, and woodland) near feeding areas. It may dwell up to
treeline The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snowp ...
, as long as the land is well-drained and shelter is available. The size and distribution of its burrow systems depend on the type of
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...
present; in areas with loose soil, it selects sites with supporting structures, such as tree roots or shrubs to prevent burrow collapse. Warrens tend to be larger and have more interconnected tunnels in areas with
chalk Chalk is a soft, white, porous, sedimentary carbonate rock. It is a form of limestone composed of the mineral calcite and originally formed deep under the sea by the compression of microscopic plankton that had settled to the sea floor. Chalk ...
than those in sand. In large coniferous plantations, the species only occurs on peripheral areas and along fire breaks and rides.


Diet

The European rabbit eats a wide variety of herbage, especially
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es, favouring the young, succulent leaves and shoots of the most nutritious species, particularly
fescue ''Festuca'' (fescue) is a genus of flowering plants belonging to the grass family Poaceae (subfamily Pooideae). They are evergreen or herbaceous perennial tufted grasses with a height range of and a cosmopolitan distribution, occurring on every ...
s. In mixed cultivated areas,
winter wheat Winter wheat (usually ''Triticum aestivum'') are strains of wheat that are planted in the autumn to germinate and develop into young plants that remain in the vegetative phase during the winter and resume growth in early spring. Classification ...
is preferred over
maize Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. Th ...
and
dicotyledon The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...
s. During the summer, the European rabbit feeds on the shortest, and therefore less nutritious grass swards, thus indicating that grazing grounds are selected through antipredator considerations rather than maximising food intake. In times of scarcity, the rabbit increases its food intake, selecting the parts of the plant with the highest
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
content. Hungry rabbits in winter may resort to eating
tree bark Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and cons ...
.
Blackberries The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus''. The taxonomy of ...
are also eaten, and captive-bred European rabbits have been fed on
fodder Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food g ...
consisting of
furze ''Ulex'' (commonly known as gorse, furze, or whin) is a genus of flowering plants in the family Fabaceae. The genus comprises about 20 species of thorny evergreen shrubs in the subfamily Faboideae of the pea family Fabaceae. The species are na ...
and
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s, which can lead to considerable weight gain. The European rabbit is a less fussy eater than the brown hare; when eating
root vegetable Root vegetables are underground plant parts eaten by humans as food. Although botany distinguishes true roots (such as taproots and tuberous roots) from non-roots (such as bulbs, corms, rhizomes, and tubers, although some contain both hypocotyl a ...
s, the rabbit eats them whole, while the hare tends to leave the peel. Depending on the body's fat and protein reserves, the species can survive without food in winter for about 2–8 days. Although
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, cases are known of rabbits eating
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...
s. Like other leporids, the European rabbit produces soft, mucus-covered faecal pellets, which are ingested directly from the
anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, d ...
. The soft pellets are produced posterior to the colon in the hind gut soon after the excretion of hard pellets and the stomach begins to fill with newly grazed food. The soft pellets are filled with
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
-rich
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...
, and pass down to the rectum in glossy clusters. The rabbit swallows them whole, without perforating the enveloping membrane.


Predators

The European rabbit is prey to many different predatory species.
Fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
es,
dingo The dingo (''Canis familiaris'', ''Canis familiaris dingo'', ''Canis dingo'', or ''Canis lupus dingo'') is an ancient (Basal (phylogenetics), basal) lineage of dog found in Australia (continent), Australia. Its taxonomic classification is de ...
es,
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
,
lynx A lynx is a type of wild cat. Lynx may also refer to: Astronomy * Lynx (constellation) * Lynx (Chinese astronomy) * Lynx X-ray Observatory, a NASA-funded mission concept for a next-generation X-ray space observatory Places Canada * Lynx, Ontar ...
es,
wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for "gluttony, glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is ...
s, and
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Do ...
s kill both adult and young rabbits by stalking and surprising them in the open, but relatively few rabbits are caught this way, as they can quickly rush back to cover with a burst of speed. Further, evidence from a study in Spain suggests they may avoid areas where the recent scat of predators which have eaten rabbit is detected. Both foxes and
badger Badgers are short-legged omnivores in the family Mustelidae (which also includes the otters, wolverines, martens, minks, polecats, weasels, and ferrets). Badgers are a polyphyletic rather than a natural taxonomic grouping, being united b ...
s dig out kittens from shallow burrows, with the latter predators being too slow to catch adult rabbits. Both
wild Wild, wild, wilds or wild may refer to: Common meanings * Wild animal * Wilderness, a wild natural environment * Wildness, the quality of being wild or untamed Art, media and entertainment Film and television * ''Wild'' (2014 film), a 2014 A ...
and
domestic cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members o ...
s can stalk and leap upon rabbits, particularly young specimens leaving their burrows for the first time. Wildcats take rabbits according to availability; in eastern Scotland, where rabbits are abundant, they can make up over 90% of the wildcats' diet. Most domestic cats are incapable of killing healthy, full-grown adults, but will take weak and diseased ones. Does can be fiercely protective of their kits, having been observed to chase away large cats and
mustelid The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in ...
s, including
ferret The ferret (''Mustela furo'') is a small, Domestication, domesticated species belonging to the family Mustelidae. The ferret is most likely a domesticated form of the wild European polecat (''Mustela putorius''), evidenced by their Hybrid (biol ...
s,
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
s, and
weasel Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets and European mink. Members of this genus are small, active predators, with long and slender bo ...
s. However, rabbits typically run from mustelids, and may fear them innately. Cases are known of rabbits becoming paralysed with fear and dying when pursued by stoats or weasels, even when rescued unharmed. The European rabbit makes up 85% of the
polecat Polecat is a common name for several mustelid species in the order Carnivora and subfamilies Ictonychinae and Mustelinae. Polecats do not form a single taxonomic rank (i.e. clade). The name is applied to several species with broad similarities t ...
's diet, and its availability is important to the success of breeding female
mink Mink are dark-colored, semiaquatic, carnivorous mammals of the genera ''Neogale'' and '' Mustela'' and part of the family Mustelidae, which also includes weasels, otters, and ferrets. There are two extant species referred to as "mink": the A ...
.
Brown rat The brown rat (''Rattus norvegicus''), also known as the common rat, street rat, sewer rat, wharf rat, Hanover rat, Norway rat, Norwegian rat and Parisian rat, is a widespread species of common rat. One of the largest muroids, it is a brown o ...
s can be a serious threat to kittens, as they will reside in rabbit burrows during the summer, and attack them in groups. Although many
birds of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators ...
are capable of killing rabbits, few are strong enough to carry them. Large species, such as
golden Golden means made of, or relating to gold. Golden may also refer to: Places United Kingdom *Golden, in the parish of Probus, Cornwall * Golden Cap, Dorset *Golden Square, Soho, London *Golden Valley, a valley on the River Frome in Gloucestershi ...
and
sea eagle A sea eagle or fish eagle (also called erne or ern, mostly in reference to the white-tailed eagle) is any of the birds of prey in the genus ''Haliaeetus'' in the bird of prey family Accipitridae. Taxonomy and evolution The genus ''Haliaeetus'' ...
s, may carry rabbits back to their nests, while small eagles,
buzzard Buzzard is the common name of several species of birds of prey. ''Buteo'' species * Archer's buzzard (''Buteo archeri'') * Augur buzzard (''Buteo augur'') * Broad-winged hawk (''Buteo platypterus'') * Common buzzard (''Buteo buteo'') * Eastern ...
s, and harriers struggle to do so.
Hawk Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
s and
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s typically only carry off very small kits.


Diseases and parasites

The European rabbit is the only species fatally attacked by
myxomatosis Myxomatosis is a disease caused by ''Myxoma virus'', a poxvirus in the genus ''Leporipoxvirus''. The natural hosts are tapeti (''Sylvilagus brasiliensis'') in South and Central America, and brush rabbits (''Sylvilagus bachmani'') in North Ame ...
. The most lethal strain has a five-day incubation period, after which the eyelids swell, with the inflammation quickly spreading to the base of the ears, the forehead, and nose. At the same time, the anal and genital areas also swell. During the last stages of the disease, the swellings discharge a fluid rich in viral material, with death usually following on the 11th-12th day of infection. In Britain, the primary carrier of myxomatosis is the flea ''
Spilopsyllus cuniculi ''Spilopsyllus cuniculi'', the rabbit flea, is a species of flea in the family Pulicidae. It is an external parasite of rabbits and hares and is occasionally found on cats and dogs and also certain seabirds that nest in burrows. It can act as a ...
'', while in Australia it is
mosquito Mosquitoes (or mosquitos) are members of a group of almost 3,600 species of small flies within the family Culicidae (from the Latin ''culex'' meaning " gnat"). The word "mosquito" (formed by ''mosca'' and diminutive ''-ito'') is Spanish for "li ...
es.
Rabbit haemorrhagic disease Rabbit hemorrhagic disease (RHD), also known as viral hemorrhagic disease (VHD), is a highly infectious and lethal form of viral hepatitis that affects European rabbits. Some viral strains also affect hares and cottontail rabbits. Mortality rate ...
(RHD), also known as viral haemorrhagic disease or rabbit calicivirus disease in Australia, is specific to the European rabbit, and causes lesions of acute necrotising
hepatitis Hepatitis is inflammation of the liver tissue. Some people or animals with hepatitis have no symptoms, whereas others develop yellow discoloration of the skin and whites of the eyes (jaundice), poor appetite, vomiting, tiredness, abdominal pa ...
, disseminated intravascular coagulation, and haemorrhaging, mainly in the lungs. Susceptible specimens may die within 30 hours of infection. Most rabbits in the UK are immune to RHD, due to exposure to a weaker strain.


Human relationship with rabbits

Recent research has shown that all European rabbits carry common genetic markers and descend from one of two maternal lines. These lines originated between 12,000 and 6.5 million years ago when glaciers isolated two herds, one on the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
and the other in
Southern France Southern France, also known as the South of France or colloquially in French language, French as , is a defined geographical area consisting of the regions of France that border the Atlantic Ocean south of the Marais Poitevin,Louis Papy, ''Le midi ...
. Humans likely began hunting rabbits as a food source, but further research needs to be done to verify this. Little comprehensive evidence of the relationship of humans with European rabbits is documented until the medieval period. Humans' relationship with the European rabbit was first recorded by the Phoenicians prior to 1000 BC, when they termed the Iberian Peninsula ''i-Shaphan-ím'' (literally, the land of the
hyraxes Hyraxes (), also called dassies, are small, thickset, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Typically, they measure between long and weigh between . They are superficially simil ...
). This phrase closely resembles related
modern Hebrew Modern Hebrew ( he, עברית חדשה, ''ʿivrít ḥadašá ', , '' lit.'' "Modern Hebrew" or "New Hebrew"), also known as Israeli Hebrew or Israeli, and generally referred to by speakers simply as Hebrew ( ), is the standard form of the He ...
: ''I'' (אי) meaning island and ''shafan'' (שפן) meaning hyrax, plural ''shfaním'' (שפנים). Phoenicians called the local rabbits 'hyraxes' because rabbits resemble hyraxes in some ways, and hyraxes are native to
Phoenicia Phoenicia () was an ancient thalassocratic civilization originating in the Levant region of the eastern Mediterranean, primarily located in modern Lebanon. The territory of the Phoenician city-states extended and shrank throughout their histor ...
, unlike rabbits. Hyraxes, like rabbits, are not rodents. One theory states that the Romans converted the phrase ''i-Shaphan-ím'', with influence from the Greek ''Spania'', to its Latin form, ''
Hispania Hispania ( la, Hispānia , ; nearly identically pronounced in Spanish, Portuguese, Catalan, and Italian) was the Roman name for the Iberian Peninsula and its provinces. Under the Roman Republic, Hispania was divided into two provinces: Hispania ...
'', which evolved in all the Iberian languages - into Castilian ''España'', Portuguese ''Espanha'', Catalan ''Espanya'' (English "Spain"), and such other variations in modern languages. Different views have been voiced on the precise meaning of ''shafan'', but the balance of opinion appears to indicate that the hyrax is indeed the intended meaning. Like the Phoenicians, neither the later
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
nor
Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of ancient Rome *''Epistle to the Romans'', shortened to ''Romans'', a letter ...
colonizers had a specific name for the rabbit, because the species is not native to
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
and
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
(though it is present there nowadays). They commonly called it "small hare" and "small digging hare", in contrast to the
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
, which is larger and doesn't make burrows.
Catullus Gaius Valerius Catullus (; 84 - 54 BCE), often referred to simply as Catullus (, ), was a Latin poet of the late Roman Republic who wrote chiefly in the neoteric style of poetry, focusing on personal life rather than classical heroes. His s ...
used the name ''cuniculus'' (a latinization of the Iberian word ''kiniklos'' and the etymological origin of the Castilian name ''conejo'', Portuguese ''coelho'' and Catalan ''conill'', and the old English name, ''coney''), and referenced its abundance in
Celtiberia The Celtiberians were a group of Celts and Celticized peoples inhabiting an area in the central-northeastern Iberian Peninsula during the final centuries BCE. They were explicitly mentioned as being Celts by several classic authors (e.g. Strab ...
by calling this region ''cuniculosa'', i.e. rabbit-ridden.López Seoane, V. (1861) ''Fáuna Mastológica de Galicia, ó historia natural de los mamíferos de este antiguo Reino, aplicada á la medicina, á la agricultura, á la industria, á las artes y al comercio.'' Manuel Mirás, 544 pages.Blázquez, J.M. (1975) ''La romanización.''
Ediciones Akal Ediciones Akal is a Spanish publisher founded in Madrid in 1972 by Ramón Akal González. It consists of a catalogue of three thousand works in forty collections which includes fields like Humanities, classic texts, modern literature, etc., and a ...
, 437 pages.
The European rabbit is the only rabbit species that has been domesticated and all 305 global rabbit breeds— from
Netherland Dwarf The Netherland Dwarf is a breed of domestic rabbit that originated in the Netherlands. Weighing , the Netherland Dwarf is one of the smallest rabbit breeds. Its popularity as a pet or show rabbit may stem from its neotenic appearance. The Net ...
to
Flemish Giant Not to be confused with the Continental Giant rabbit The Flemish Giant rabbit is the largest breed of domestic rabbit (''Oryctolagus cuniculus domesticus''). Flemish Giants are historically a utility breed used for their fur and meat. They are o ...
— are descendants of the European rabbit. Rabbits are an example of an animal that can be treated as a food, a pet, or a pest by different members of the same culture. In some urban areas, infestations of feral European rabbits (descended from pets) have become a problem.
Helsinki Helsinki ( or ; ; sv, Helsingfors, ) is the Capital city, capital, primate city, primate, and List of cities and towns in Finland, most populous city of Finland. Located on the shore of the Gulf of Finland, it is the seat of the region of U ...
, for example, host to one of the northernmost populations of the species, had an estimated 2,500 European rabbits at the end of 2006, doubling to 5,000 by autumn 2007. In Iceland, populations of ''O. cuniculus'' are found in urban Reykjavik as well as in the
Vestmannaeyjar Vestmannaeyjar (, sometimes anglicized as Westman Islands) is a municipality and archipelago off the south coast of Iceland. The largest island, Heimaey, has a population of 4,414, most of whom live in the archipelago's main town, Vestmannaeyja ...
archipelago. In Finland, the introduced European rabbit vies with the native lagomorphs: the
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
and the
mountain hare The mountain hare (''Lepus timidus''), also known as blue hare, tundra hare, variable hare, white hare, snow hare, alpine hare, and Irish hare, is a Palearctic hare that is largely adapted to polar and mountainous habitats. Evolution The mountai ...
.


As an introduced species

The European rabbit has been introduced as an
exotic species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
into several environments, often with harmful results to vegetation and local wildlife, making it an
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
. The first known mention of the rabbit as an invasive species (and possibly the first documented instance of an invasive species ever) was made in regard to the introduction of the rabbit to the
Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
after the Roman conquest of the first century BCE. According to both
Strabo Strabo''Strabo'' (meaning "squinty", as in strabismus) was a term employed by the Romans for anyone whose eyes were distorted or deformed. The father of Pompey was called "Pompeius Strabo". A native of Sicily so clear-sighted that he could see ...
and
Pliny the Elder Gaius Plinius Secundus (AD 23/2479), called Pliny the Elder (), was a Roman author, naturalist and natural philosopher, and naval and army commander of the early Roman Empire, and a friend of the emperor Vespasian. He wrote the encyclopedic '' ...
, the multiplying rabbits caused famines by destroying crop yields and even collapsed trees and houses with their burrowing. The inhabitants petitioned
Augustus Caesar Augustus (born Gaius Octavius; 23 September 63 BC – 19 August AD 14), also known as Octavian, was the first Roman emperor; he reigned from 27 BC until his death in AD 14. He is known for being the founder of the Roman Pri ...
for help, who sent troops to curb the rabbit population with the help of
ferret The ferret (''Mustela furo'') is a small, Domestication, domesticated species belonging to the family Mustelidae. The ferret is most likely a domesticated form of the wild European polecat (''Mustela putorius''), evidenced by their Hybrid (biol ...
s. Other locations where the European rabbit was introduced include
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
; the Hawaiian Islands of
Laysan Island Laysan (; haw, italics=no, Kauō ), located northwest of Honolulu at , is one of the Northwestern Hawaiian Islands. It comprises one land mass of , about in size. It is an atoll of sorts, although the land completely surrounds a shallow ...
(in 1903) and
Lisianski Island Lisianski Island ( Hawaiian: ''Papa‘āpoho'') is one of the Northwestern Hawaiian Islands, with a land area of and a maximum elevation of above sea level. It is a low, flat sand and coral island about northwest of Honolulu, Hawaii. The is ...
; Oceania's
Macquarie Island Macquarie Island is an island in the southwestern Pacific Ocean, about halfway between New Zealand and Antarctica. Regionally part of Oceania and politically a part of Tasmania, Australia, since 1900, it became a Tasmanian State Reserve in 197 ...
; Washington's Smith Island and
San Juan Island San Juan Island is the second-largest and most populous of the San Juan Islands in northwestern Washington, United States. It has a land area of 142.59 km2 (55.053 sq mi) and a population of 6,822 as of the 2000 census. Washington State Fe ...
(around 1900 and later spreading to the other San Juan Islands); several islands off the coast of
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
(including
Robben Island Robben Island ( af, Robbeneiland) is an island in Table Bay, 6.9 kilometres (4.3 mi) west of the coast of Bloubergstrand, north of Cape Town, South Africa. It takes its name from the Dutch word for seals (''robben''), hence the Dutch/Afrik ...
); and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
. The two accounts over the introduction of rabbits in
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
are conflicting. One holds that the species was brought there in the early 20th century by Austrian nobleman
Graf (feminine: ) is a historical title of the German nobility, usually translated as "count". Considered to be intermediate among noble ranks, the title is often treated as equivalent to the British title of "earl" (whose female version is "coun ...
Malokhovsky, who released them on his estate near the
Khadzhibey Estuary Khadzhibey Estuary, or Khadzhibeyskyi Liman ( uk, Хаджибейський лиман, tr, Haci Bey limanı), is an estuary of the north-western part of the Black Sea, located on the north-west from the City of Odessa. It is named after the for ...
, while another holds that rabbits were first brought to
Kherson Kherson (, ) is a port city of Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers appr ...
from
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
in 1894-1895 by landowner Pinkovsky.


In the British Isles

The European rabbit is widespread in both Great Britain,
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
, and most islands, except for
Isles of Scilly The Isles of Scilly (; kw, Syllan, ', or ) is an archipelago off the southwestern tip of Cornwall, England. One of the islands, St Agnes, is the most southerly point in Britain, being over further south than the most southerly point of the ...
,
Rùm Rùm (), a Scottish Gaelic name often anglicised to Rum (), is one of the Small Isles of the Inner Hebrides, in the district of Lochaber, Scotland. For much of the 20th century the name became Rhum, a spelling invented by the former owner, Sir ...
,
Tiree Tiree (; gd, Tiriodh, ) is the most westerly island in the Inner Hebrides of Scotland. The low-lying island, southwest of Coll, has an area of and a population of around 650. The land is highly fertile, and crofting, alongside tourism, and ...
, and some small Scottish islands, such as Gunna, Sanday, and most of the
Treshnish Isles The Treshnish Isles are an archipelago of small islands and skerries, lying west of the Isle of Mull, in Scotland. They are part of the Inner Hebrides. Trips to the Treshnish Isles operate from Ulva Ferry, Tobermory, Ardnamurchan and Tiree. Geo ...
. It was likely first brought to Britain by the
Normans The Normans (Norman language, Norman: ''Normaunds''; french: Normands; la, Nortmanni/Normanni) were a population arising in the medieval Duchy of Normandy from the intermingling between Norsemen, Norse Viking settlers and indigenous West Fran ...
after the 1066 conquest of England, as no pre-Norman British allusions to the animal have been found. The rabbit was nonetheless scarce or absent throughout most of England a short time afterwards, as warrens are not mentioned in the
Domesday Book Domesday Book () – the Middle English spelling of "Doomsday Book" – is a manuscript record of the "Great Survey" of much of England and parts of Wales completed in 1086 by order of King William I, known as William the Conqueror. The manusc ...
or any other 11th-century documents. Rabbits became well known, but not necessarily accepted members of British fauna between the 12th and 13th centuries, with the first real evidence of their presence consisting of a number of bones from the
midden A midden (also kitchen midden or shell heap) is an old dump for domestic waste which may consist of animal bone, human excrement, botanical material, mollusc shells, potsherds, lithics (especially debitage), and other artifacts and ecofact ...
of
Rayleigh Castle Rayleigh Castle (also known as Rayleigh Mount) was a masonry and timber castle built in the town of Rayleigh in Essex, England in the 11th century shortly after the Norman conquest. All that exists today are the earthwork remains of its large ...
, which was occupied from the 11th-13th centuries. The first references to rabbits in Ireland occur roughly at the same time as English ones, thus indicating another Norman introduction. They had become plentiful, probably at a local level, by the 13th century, as indicated by an inquisition of
Lundy Island Lundy is an English island in the Bristol Channel. It was a micronation from 1925–1969. It forms part of the district of Torridge in the county of Devon. About long and wide, Lundy has had a long and turbulent history, frequently chang ...
made in 1274 describing how 2,000 rabbits were caught annually. Subsequent allusions in official documents became more frequent, with the species later becoming an important food item in feasts. Increases in truly wild populations occurred slowly, primarily in the coastal areas and lowland heaths of Breckland and
Norfolk Norfolk () is a ceremonial and non-metropolitan county in East Anglia in England. It borders Lincolnshire to the north-west, Cambridgeshire to the west and south-west, and Suffolk to the south. Its northern and eastern boundaries are the No ...
. Notable population increases occurred after 1750, when changes in agricultural practices created favourable habitat, and increasing interest in game management resulted in intensive predator control campaigns. Although now common in the Scottish lowlands, the species was little known in Scotland before the 19th century. Until then, it was confined to portions of the
Edinburgh Edinburgh ( ; gd, Dùn Èideann ) is the capital city of Scotland and one of its 32 Council areas of Scotland, council areas. Historically part of the county of Midlothian (interchangeably Edinburghshire before 1921), it is located in Lothian ...
district at least as far back as the 16th century, certain islands and the coastal
sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, fl ...
s of the Scottish mainland. Although unknown in
Caithness Caithness ( gd, Gallaibh ; sco, Caitnes; non, Katanes) is a historic county, registration county and lieutenancy area of Scotland. Caithness has a land boundary with the historic county of Sutherland to the west and is otherwise bounded by ...
in 1743, the species became well established there by 1793.
Myxomatosis Myxomatosis is a disease caused by ''Myxoma virus'', a poxvirus in the genus ''Leporipoxvirus''. The natural hosts are tapeti (''Sylvilagus brasiliensis'') in South and Central America, and brush rabbits (''Sylvilagus bachmani'') in North Ame ...
entered Britain from France in 1953, and reached Ireland by 1954, prompting the
RSPB The Royal Society for the Protection of Birds (RSPB) is a Charitable_organization#United_Kingdom, charitable organisation registered in Charity Commission for England and Wales, England and Wales and in Office of the Scottish Charity Regulator, ...
to set up "mercy squads" meant to euthanise myxomatous rabbits. Major myxomatosis outbreaks still occur in Britain, peaking twice annually in during spring and especially in the late summer or autumn periods, though
immunity Immunity may refer to: Medicine * Immunity (medical), resistance of an organism to infection or disease * ''Immunity'' (journal), a scientific journal published by Cell Press Biology * Immune system Engineering * Radiofrequence immunity desc ...
has reduced the
mortality rate Mortality rate, or death rate, is a measure of the number of deaths (in general, or due to a specific cause) in a particular population, scaled to the size of that population, per unit of time. Mortality rate is typically expressed in units of de ...
from 99% to 5-33%. Between 1996 and 2018, rabbit numbers fell by 88% in the east Midlands, 83% in Scotland, and 43% across the whole of the UK. Numbers are still falling (in 2021). Pip Mountjoy, Shifting Sands project manager at
Natural England Natural England is a non-departmental public body in the United Kingdom sponsored by the Department for Environment, Food and Rural Affairs. It is responsible for ensuring that England's natural environment, including its land, flora and fauna, ...
said, "They (rabbits) are actually an endangered species in their native region on the Iberian peninsula. It's surprising for people that rabbits are important in some ecosystems. We think of them as a pest but in Britain they are a keystone species – they act as landscape managers and a lot of other species rely on them." The Shifting Sands project aims to encourage landowners to create safe habitats for rabbits, consisting of piles of branches placed near existing rabbit warrens. Species that depend in rabbits' grazing habits include purple milk vetch, rare spring sedge, spring speedwell, prostrate perennial knawel, caterpillars of the lunar yellow underwing moth, stone curlew, and the
large blue butterfly The large blue (''Phengaris arion'') is a species of butterfly in the family Lycaenidae. The species was first defined in 1758 and first recorded in Great Britain, Britain in 1795. In 1979 the species became mostly extinct in Britain but has be ...
.


In Australia

Twenty-four specimens of the European rabbit were introduced to Australia in 1859 by estate owner
Thomas Austin Thomas Austin may refer to: *Thomas Austin (pastoralist) (1815–1871), English settler who introduced rabbits into Australia * Thomas Austin (American football) (born 1986), former American football center * Thomas Austin (cricketer) (1857–1941) ...
in
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
. Their descendants multiplied and spread throughout the country due to the lack of natural predators, a conducive habitat (provided by widespread farming), and the mild Australian winters that allowed year-round breeding. Australia's native equivalent, the bilby, was quickly pushed out by the invasive rabbit. (The bilbies are endangered, but are now making a comeback due to government protection.) Between 1901 and 1907, Australia built an immense "
rabbit-proof fence The State Barrier Fence of Western Australia, formerly known as the Rabbit-Proof Fence, the State Vermin Fence, and the Emu Fence, is a pest-exclusion fence constructed between 1901 and 1907 to keep rabbits, and other agricultural pests from th ...
" to halt the westward expansion of the infestation. The European rabbit, however, can not only jump very high, but also burrow underground, making fencing essentially futile. During the 1950s, the intentional introduction of a virus that causes myxomatosis provided some relief in Australia, but not in New Zealand, where the insect
vector Vector most often refers to: *Euclidean vector, a quantity with a magnitude and a direction *Vector (epidemiology), an agent that carries and transmits an infectious pathogen into another living organism Vector may also refer to: Mathematic ...
s necessary for the spread of the disease were not present. Myxomatosis can also infect pet rabbits (the same species). Today's remaining feral rabbits in Australia are largely immune to myxomatosis. A second deadly rabbit virus,
rabbit hemorrhagic disease Rabbit hemorrhagic disease (RHD), also known as viral hemorrhagic disease (VHD), is a highly infectious and lethal form of viral hepatitis that affects European rabbits. Some viral strains also affect hares and cottontail rabbits. Mortality rate ...
(RHD), has been cleared in Australia as a
biological control Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also invo ...
agent and has already killed millions of the European rabbits there. RHD was also introduced—illegally— in New Zealand with less success due to improper timing.


In Chile

The exact date on which the European rabbit was introduced into
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
is unknown, though the first references to it occur during the mid-18th century. By the 19th century, several authors referred to the presence of both rabbits and rabbit hutches in central Chile. The importation and breeding of rabbits was encouraged by the state, as rabbits were seen as cheap sources of food for peasants. Whether or not their escape into the wild was intentional is unknown, but warnings over the dangers of feral rabbits were raised during the early 20th century, and the species had propagated dramatically by the late 1920s in central Chile,
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla G ...
, and the
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
. In the 1930s, the state sought to tackle the rabbit problem by banning fox hunting, though it was later discovered that indigenous
South American fox The South American foxes (''Lycalopex''), commonly called ''raposa'' in Portuguese language, Portuguese, or ''zorro'' in Spanish language, Spanish, are a genus from South America of the subfamily Caninae. Despite their name, they are not Vulpes, ...
es rarely preyed on rabbits, preferring native species. In modern times, the European rabbit problem has not been resolved definitively, though a deliberate outbreak of myxomatosis in Tierra del Fuego successfully reduced local rabbit populations. The species remains a problem in central Chile and on Juan Fernández, despite international financing.


Domesticated rabbits

The European rabbit is the only rabbit to be widely
domesticated Domestication is a sustained multi-generational relationship in which humans assume a significant degree of control over the reproduction and care of another group of organisms to secure a more predictable supply of resources from that group. A ...
, for food or as a
pet A pet, or companion animal, is an animal kept primarily for a person's company or entertainment rather than as a working animal, livestock, or a laboratory animal. Popular pets are often considered to have attractive appearances, intelligence, ...
. It was first widely kept in
ancient Rome In modern historiography, ancient Rome refers to Roman civilisation from the founding of the city of Rome in the 8th century BC to the collapse of the Western Roman Empire in the 5th century AD. It encompasses the Roman Kingdom (753–509 B ...
, where fetal rabbits were known as ''
laurices The term laurices refers to the fetus of the rabbit (''Oryctolagus cuniculus'') prepared without evisceration and consumed as a table delicacy. The word is the plural of the Latin word laurex (variant ''laurix'', n. masc., pl. ''laurices''; Englis ...
'' and considered a delicacy, and has been refined into a wide variety of breeds during and since the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire a ...
. Domesticated rabbits have mostly been bred to be much larger than wild rabbits, though
selective breeding Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant mal ...
has produced a range of sizes from "dwarf" to "giant", which are kept as food animals and pets across the world. They have as much color variation among themselves as other livestock and pet animals. Their fur is prized for its softness; today,
Angora rabbits The Angora rabbit ( tr, Ankara tavşanı), which is one of the oldest types of domestic rabbit, is bred for the long fibers of its coat, known as ''Angora wool'', which are gathered by Sheep shearing, shearing, combing or plucking (hair removal), ...
are raised for their long, soft fur, which is often spun into yarn. Other breeds are raised for the fur industry, particularly the Rex, which has a smooth,
velvet Weave details visible on a purple-colored velvet fabric Velvet is a type of woven tufted fabric in which the cut threads are evenly distributed, with a short pile, giving it a distinctive soft feel. By extension, the word ''velvety'' means ...
-like coat and occurs in a wide variety of colors and sizes.


Meat and fur

In the United Kingdom, rabbit was a popular food source for the poorer classes. Among wild rabbits, those native to Spain were reputed to have the highest meat quality, followed by those in the
Ardennes The Ardennes (french: Ardenne ; nl, Ardennen ; german: Ardennen; wa, Årdene ; lb, Ardennen ), also known as the Ardennes Forest or Forest of Ardennes, is a region of extensive forests, rough terrain, rolling hills and ridges primarily in Be ...
. As rabbits hold very little fat, they were hardly ever roasted, being instead boiled, fried, or stewed. The pelt of the rabbit is heavier and more durable than the hare's.Petersen, Marcus (1914),
The fur traders, and fur bearing animals
', Buffalo : Hammond Press, p. 291
Marshall Marshall may refer to: Places Australia * Marshall, Victoria, a suburb of Geelong, Victoria Canada * Marshall, Saskatchewan * The Marshall, a mountain in British Columbia Liberia * Marshall, Liberia Marshall Islands * Marshall Islands, an i ...
calculated that the value of the skin in proportion to the carcass was greater than that of the
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
and ox. Its fur is primarily used for
felt Felt is a textile material that is produced by matting, condensing and pressing fibers together. Felt can be made of natural fibers such as wool or animal fur, or from synthetic fibers such as petroleum-based acrylic or acrylonitrile or wood ...
ing or
hat A hat is a head covering which is worn for various reasons, including protection against weather conditions, ceremonial reasons such as university graduation, religious reasons, safety, or as a fashion accessory. Hats which incorporate mecha ...
s. It is also dyed or clipped, and sold as imitations of more valuable furbearers, such as
fur seal Fur seals are any of nine species of pinnipeds belonging to the subfamily Arctocephalinae in the family '' Otariidae''. They are much more closely related to sea lions than true seals, and share with them external ears (pinnae), relatively lon ...
. Although cheap and easily acquired, rabbit fur has little durability.


Conservation status

Though the European rabbit thrives in many of the locations where it was introduced, in its native Iberia, populations are dwindling. In 2005, the Portuguese
Institute for Nature Conservation and Forests The Institute for Nature Conservation and Forests ( Portuguese: ''Instituto da Conservação da Natureza e das Florestas, I.P.''), ICNF, is the responsible governmental body for the nature and forest policies, including the management of Protected ...
classified ''O. cuniculus'' in Portugal as "near threatened", while in 2006, Spanish authorities (SECEM) reclassified it in Spain as "vulnerable". In 2018, the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
reclassified ''O. cuniculus'' in Spain, Portugal, and France as "endangered", due to the extent of recent declines. However, worldwide, the species is
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
.


See also

*
Cuniculture Cuniculture is the agricultural practice of breeding and raising domestic rabbits as livestock for their meat, fur, or wool. Cuniculture is also employed by rabbit fanciers and hobbyists in the development and betterment of rabbit breeds and the ...
, on the practice of breeding and raising the domesticated version of the European rabbit * List of breeds of the domesticated version of the European rabbit


References


Bibliography

* * * * *


External links

* * * View th
rabbit genome
in
Ensembl Ensembl genome database project is a scientific project at the European Bioinformatics Institute, which provides a centralized resource for geneticists, molecular biologists and other researchers studying the genomes of our own species and other v ...
* {{Authority control Leporidae Mammals described in 1758 Fauna of Gibraltar Mammals of Europe Mammals of North Africa Taxa named by Carl Linnaeus