The Orinoco crocodile (''Crocodylus intermedius'') is a
critically endangered crocodile
Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...
. Its population is very small, and they can only be found in the Orinoco river basin in
Colombia
Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
and
Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. Extensively hunted for their skins in the 19th and 20th centuries, it is one of the most endangered species of crocodiles. It is a very large species of
crocodilian
Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
; males have been reported up to in the past, but such sizes do not exist today, being a more widely accepted maximum size. A large male today may attain in length and can weigh up to , while females are substantially smaller with the largest likely to weigh around .
Sexual dimorphism
Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
is not as profound as in other crocodilian species. The coloration is light even in adults.
The biology of the Orinoco crocodile is poorly documented in the wild, mostly due its small population. It is thought to have a more
piscivorous
A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evoluti ...
diet with an
opportunistic
Opportunism is the practice of taking advantage of circumstances – with little regard for principles or with what the consequences are for others. Opportunist actions are expedient actions guided primarily by self-interested motives. The term ...
nature, resulting in
generalist
A generalist is a person with a wide array of knowledge on a variety of subjects, useful or not. It may also refer to:
Occupations
* a physician who provides general health care, as opposed to a medical specialist; see also:
** General pract ...
predatory behavior. The Orinoco crocodile is an
apex predator
An apex predator, also known as a top predator, is a predator at the top of a food chain, without natural predators of its own.
Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the highest trophic lev ...
, and will take the opportunity to prey on a variety of
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s,
mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s, and
reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s, including
caiman
A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South America fro ...
s on occasion. The prey base is mostly made up of large predatory
fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
, challenging the general view by locals complaining about crocodiles hunting local fish to very low numbers. Despite its large size, the Orinoco crocodile rarely poses a threat to humans, despite several reports. Reproduction takes place in the
dry season
The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The te ...
when the water level is lower. Orinoco crocodiles are hole nesters, just like the related
Nile
The Nile, , Bohairic , lg, Kiira , Nobiin language, Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered ...
,
freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
, and
saltwater crocodile
The saltwater crocodile (''Crocodylus porosus'') is a crocodilian native to saltwater habitats and brackish wetlands from India's east coast across Southeast Asia and the Sundaic region to northern Australia and Micronesia. It has been listed ...
s, and they dig holes in the sand to lay their
egg
An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s. The females guard the
nest
A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
s and young for several years.
Taxonomy
The
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Crocodylus
''Crocodylus'' is a genus of true crocodiles in the family Crocodylidae.
Taxonomy
The generic name, ''Crocodylus'', was proposed by Josephus Nicolaus Laurenti in 1768. ''Crocodylus'' contains 13–14 extant (living) species and 5 extinct species ...
'' likely originated in Africa and
radiated outwards towards Southeast Asia and the Americas,
although an Australia/Asia origin has also been considered.
Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
evidence supports ''Crocodylus'' diverging from its closest recent relative, the extinct ''
Voay
''Voay'' is an extinct genus of crocodile from Madagascar that lived during the Late Pleistocene to Holocene, containing only one species, ''V. robustus''. Numerous subfossils have been found, including complete skulls, noted for their distinctiv ...
'' of
Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
, around 25 million years ago, near the
Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
/
Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
boundary.
Below is a
cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
based on a 2018
tip dating Tip dating is a technique used in molecular dating that allows the inference of time-calibrated phylogenetic trees. Its defining feature is that it uses the ages of the samples to provide time information for the analysis, in contrast with traditio ...
study by Lee & Yates simultaneously using
morphological, molecular (
DNA sequencing
DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
), and
stratigraphic
Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks.
Stratigraphy has three related subfields: lithostrati ...
(
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
age) data,
as revised by the 2021 Hekkala ''et al.''
paleogenomics Paleogenomics is a field of science based on the reconstruction and analysis of genomic information in extinct species. Improved methods for the extraction of ancient DNA (aDNA) from museum artifacts, ice cores, archeological or paleontological site ...
study using DNA extracted from the extinct ''
Voay
''Voay'' is an extinct genus of crocodile from Madagascar that lived during the Late Pleistocene to Holocene, containing only one species, ''V. robustus''. Numerous subfossils have been found, including complete skulls, noted for their distinctiv ...
''.
Characteristics
![Croc inter](https://upload.wikimedia.org/wikipedia/commons/f/f2/Croc_inter.jpg)
The Orinoco crocodile can be recognised by its relatively long snout, which is narrower than that of the somewhat similar-looking
American crocodile
The American crocodile (''Crocodylus acutus'') is a species of crocodilian found in the Neotropics. It is the most widespread of the four extant species of crocodiles from the Americas, with populations present from South Florida and the coasts ...
. This species generally has a pale tan hide, though at least three coloration variations are known, with some almost completely yellowish, and some a dark brownish-gray. The skin can change colour over long periods of time; this phenomenon has been recorded in other species that can gradually change the amount of
melanin
Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino ...
in their skin. These crocodiles have dark-brown markings, which present as more pronounced bands in younger specimens and as scattered markings on mature ones.
[Ross, R.P. (1998) ''Crocodiles: Status Survey and Conservation Action Plan. Second Edition''. IUCN/SSC Crocodile Specialist Group, IUCN, Gland, Switzerland and Cambridge, UK.] One individual measuring and weighing had a bite force of .
Size
The Orinoco crocodile ranks among the
largest living reptiles, as well as the largest
predator
Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
in the Americas. It is arguably, on average, the largest
crocodilian
Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
in the Americas; while American crocodiles,
black caiman
The black caiman (''Melanosuchus niger'') is a species of large crocodilian and is the largest species of the family Alligatoridae. It is a carnivorous reptile that lives along slow-moving rivers, lakes, seasonally flooded savannas of the Amazon ...
s and the
American alligator
The American alligator (''Alligator mississippiensis''), sometimes referred to colloquially as a gator or common alligator, is a large crocodilian reptile native to the Southeastern United States. It is one of the two extant species in the g ...
may approach similar dimensions, the Orinoco crocodile may be (or had been, when in healthy numbers) slightly longer.
Given its possible maximum sizes, the Orinoco crocodile may rank as the third largest extant true crocodile, after the
saltwater crocodile
The saltwater crocodile (''Crocodylus porosus'') is a crocodilian native to saltwater habitats and brackish wetlands from India's east coast across Southeast Asia and the Sundaic region to northern Australia and Micronesia. It has been listed ...
and
Nile crocodile
The Nile crocodile (''Crocodylus niloticus'') is a large crocodilian native to freshwater habitats in Africa, where it is present in 26 countries. It is widely distributed throughout sub-Saharan Africa, occurring mostly in the central, eastern ...
(which is closely related to it, despite its substantially different range), and additionally rank 4th amongst all extant crocodilians behind the
gharial
The gharial (''Gavialis gangeticus''), also known as gavial or fish-eating crocodile, is a crocodilian in the family Gavialidae and among the longest of all living crocodilians. Mature females are long, and males . Adult males have a distinct b ...
, though there is little to suggest that Orinoco specimens in modern times can rival these species.
[ Sexual maturity for Orinoco crocodiles is obtained for females at around while that of males seems to be obtained around length, with most adult crocodiles of the species exceeding . Average length of wild adult females from 1985 to 1992 was found to be while that of males is not known to have been surveyed at that time.] If not culled by humans, mature males easily exceed and attain a length of as much as in length as they grow throughout life, perhaps weighing up to in big specimens, while females may grow over the expected size of in length and may sometimes exceed . According to Guinness Records
''Guinness World Records'', known from its inception in 1955 until 1999 as ''The Guinness Book of Records'' and in previous United States editions as ''The Guinness Book of World Records'', is a reference book published annually, listing world ...
, the average length of adults that they were able to examine was only and the largest specimen found firsthand was .[ The largest specimen historically reported was shot in 1800, and allegedly measured at , although the source is considered reliable, unfortunately, no voucher specimen is known from this.] Because of extensive hunting for their skins in the 20th century, such giants do not exist today, and modern Orinoco crocodiles have not been reported to exceed in length.[ Merchán listed the maximum total length attainable for a male as and the maximum length of a female as .
]
Distribution and habitat
This species is restricted to the Orinoco river basin in Colombia and Venezuela. They have been occasionally reported on the island of Trinidad
Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...
, but this has not been confirmed, and witnesses may have mistaken an American crocodile for the rarer species. This crocodile was once thought to have inhabited a wide range of riparian
A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...
habitats, from tropical forests
Tropical forests (a.k.a. jungle) are forested landscapes in tropical regions: ''i.e.'' land areas approximately bounded by the tropic of Cancer and Capricorn, but possibly affected by other factors such as prevailing winds.
Some tropical fores ...
to the streams of the Andes
The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
foothills. Today, this species is restricted to the Llanos savanna and associated seasonal freshwater rivers.[
]
Behavior and ecology
Hunting and diet
Little study has gone into the dietary biology of Orinoco crocodiles but from eye-witness accounts and partial studies from captivity and crocodile farms, the majority of the Orinoco crocodile's diet appears to consists of large fish. The species' relatively narrow snout is ideally suited to minimize water resistance in capturing such aquatic prey. However, as an opportunistic apex predator, virtually any animal living within its range could be considered a potential meal, such as invertebrates, reptiles, birds, and mammals. Despite having a rather elongated skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
, its base snout is wide, hinting to a generalist
A generalist is a person with a wide array of knowledge on a variety of subjects, useful or not. It may also refer to:
Occupations
* a physician who provides general health care, as opposed to a medical specialist; see also:
** General pract ...
diet. Additionally, as they age, mature males in particular appear to manifest a broadened snout, presumably due to a shift to being able to take larger prey as is seen in several other large crocodile species. It stalks both aquatic and terrestrial
Terrestrial refers to things related to land or the planet Earth.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
prey. As a large adult, terrestrial prey may include monkey
Monkey is a common name that may refer to most mammals of the infraorder Simiiformes, also known as the simians. Traditionally, all animals in the group now known as simians are counted as monkeys except the apes, which constitutes an incomple ...
s, deer
Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
, bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s, other reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s, domestic animal
This page gives a list of domesticated animals, also including a list of animals which are or may be currently undergoing the process of domestication and animals that have an extensive relationship with humans beyond simple predation. This includ ...
s, and even occasionally other large predators if the opportunity arises.[ Similar to many of the larger crocodile species, the Orinoco crocodile has also been observed catching and eating smaller species of crocodilians, such as adult common caimans and sometimes cannibalizing smaller individuals of its own kind.
Attacks on humans have been reported, but this is highly unlikely to be a common behavior today, given the very small population of the species and its relative isolation from large human settlements. Historically attacks were not unusual and on his trip to the region in 1800, natives told ]Alexander von Humboldt
Friedrich Wilhelm Heinrich Alexander von Humboldt (14 September 17696 May 1859) was a German polymath, geographer, naturalist, explorer, and proponent of Romantic philosophy and science. He was the younger brother of the Prussian minister, p ...
that two or three adult people were killed per year by Orinoco crocodiles. A small number of better documented fatal attacks were reported in the 1900s–1930s when the species was still relatively common. The only well-documented recent attack, on a fisherman in 2009, was serious but not fatal.[ A second survivor was reported in 2011 to live in La Palmita at the ]Cojedes River
The Cojedes River (Spanish ''Rio Cojedes'') is a tributary of the Orinoco River in central Venezuela. The Cojedes originates in Lara state, and flows southeast through a gap between the Cordillera de Mérida and the Cordillera de la Costa and ac ...
, but any details of this attack (including when exactly it happened) are lacking.[
]
Reproduction
When water recedes in the dry season
The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The te ...
, Orinoco crocodiles retreat to burrows they excavate into the riverbanks. The adult pair mates during the drier period of the year. Adult males attract the females by roaring. The females often outnumber the males two to one and appear to select the males with the deepest roars. In a reintroduced population, only 6 of 14 potential adult males parented over 90% of eggs laid. Usually 14 weeks after mating, the female crocodile will dig a nest and lay about 40 eggs. It is a hole-nester, as are all crocodilians, and it usually makes its nest on a sand bank. The eggs incubate under a mixture of soil and rotting vegetation for around three months. The most common predator of buried eggs are tegu lizards although the tegus are sometimes caught and killed by the mother crocodile.[ During the night the young hatch and call to their mother; she digs them out of the nest and carries them to the water, which is considerably higher at this point. Young Orinoco crocodiles are often at risk from predation by American black vultures, tegu lizards, ]anaconda
Anacondas or water boas are a group of large snakes of the genus ''Eunectes''. They are found in tropical South America. Four species are currently recognized.
Description
Although the name applies to a group of snakes, it is often used to re ...
s, caimans
A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South America fro ...
, coati
Coatis, also known as coatimundis (), are members of the family Procyonidae in the genera ''Nasua'' and ''Nasuella''. They are diurnal mammals native to South America, Central America, Mexico, and the southwestern United States. The name "c ...
s, jaguar
The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
s, and other carnivore
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
s, though these species are sometimes also caught and killed by the defending mother crocodile. Adults have no predators except for humans. Females have defended pods of juveniles for over three years, though closer to one year to independence is generally most common.[ A study of captive Orinoco crocodiles noted the aggressive behavior of adults while nesting and noted that the normally relatively docile crocodilians could not be approached while they were actively brooding. Without persecution, it is possible that Orinoco crocodiles may reach a lifespan of 70–80 years.][
]
Conservation status
The Orinoco crocodile is highly endangered due to excessive hunting for its hide. During the 1940s to the 1960s, thousands of these animals were slaughtered in the Orinoco River and the Llanos wetlands, and the species came very close to extinction
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
. The Orinoco crocodile was given protected status in the 1970s, but has yet to recover. Today, it is protected both in Colombia and Venezuela, and also included on Appendix I by CITES
CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
. In addition to hunting for its hide, more recent threats include the collection of juveniles for sale in the live animal trade, pollution
Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the ...
, and the proposal of a dam
A dam is a barrier that stops or restricts the flow of surface water or underground streams. Reservoirs created by dams not only suppress floods but also provide water for activities such as irrigation, human consumption, industrial use, a ...
in the upper Orinoco River region. Another problem is the increased population of spectacled caimans, a smaller crocodilian that can outcompete the Orinoco crocodile for fish due to its much larger population and much more accelerated breeding rates.[
It's unclear how many individuals remain in the wild, but estimates range between 250 and 1500.][Information on ''Crocodylus intermedius''.]
ARKIVE. The largest subpopulation in Venezuela is in Cojedes and Sarare
Sarare is a town in the Venezuelan state of Lara. It is the seat of the Simón Planas Municipality. The town and surroundings have about 12535 inhabitants. It is located on the shores of the Sarare River, which flows into the Orinoco
The Ori ...
, with fewer than 500 adults remaining. A number of other smaller subpopulations exist.[IUCN/SSC ''Crocodylus intermedius''.]
Crocodile Specialist Group – Status Survey and Conservation Action Plan, 2nd edition.
In November 2007, 50 individuals were held in zoo
A zoo (short for zoological garden; also called an animal park or menagerie) is a facility in which animals are kept within enclosures for public exhibition and often bred for Conservation biology, conservation purposes.
The term ''zoological g ...
s registered by Species360
Species360 (formerly International Species Information System or ISIS) founded in 1974, is an international non-profit organization that maintains an online database of wild animals under human care. , the organization serves more than 1,000 zoo ...
, of which the largest population, 35 individuals, were kept in the Dallas World Aquarium
The Dallas World Aquarium is a for-profit aquarium and zoo located in the West End Historic District of Dallas, Texas, USA. It aids conservation and education by housing many animals that are threatened or endangered as part of a cooperative br ...
. Additionally, a large number of individuals are held at captive-breeding facilities in Venezuela. Since the early 1990s, a large number of hatchlings have been released both into private ranchlands (especially in the Llanos where nature-oriented tourism is important for the local economy) and in national parks in Venezuela. While six Venezuelan captive-breeding programs continue today, many are plagued by lack of funds or staff, as well as conflicts between private and state-owned facilities.
The Orinoco crocodile became part of ''Proyecto Vida Silvestre'', a program launched in 2014 to protect 10 wildlife species of Colombia's Llanos
The Llanos (Spanish ''Los Llanos'', "The Plains"; ) is a vast tropical grassland plain situated to the east of the Andes in Colombia and Venezuela, in northwestern South America. It is an ecoregion of the tropical and subtropical grasslands, sav ...
. Thanks to that program, between May 2015 and February 2016, 41 orinoco crocodiles were reintroduced at El Tuparro National Natural Park
El Tuparro National Natural Park ( es, Parque Nacional Natural El Tuparro) is a national park located in the Vichada Department in the Orinoquía Region of Colombia. It is the only protected area in the Eastern Plains ('' Llanos Orientales'') un ...
in eastern Colombia.Endangered Colombian crocodiles get a helping hand
Fox News (15 February 2016). Retrieved on 2019-01-25.
References
External links
{{Taxonbar, from=Q272395
Crocodylidae
Reptiles of Colombia
Reptiles of Venezuela
Reptiles described in 1819
Crocodilians of South America
Apex predators