Orculella Mesopotamica Mesopotamica
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''Orculella'' is a genus of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
s, terrestrial
pulmonate Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the Middle East, the Aegean Islands (mainly Crete) and Libya ( Cyrenaica). A few species inhabit Southern Spain ( ''O. bulgarica''), Sicily ( ''O. templorum'') and northeastern Africa ( ''O. tingitana''). Species in the genera '' Schileykula'' and ''Orculella'' are indistinguishable based on shell characters only. They differ however in the genitalia, primarily in the presence (''Orculella'') or absence ('' Schileykula'') of a penial appendix.


Species and subspecies

Species and subspecies in the genus ''Orculella'' include: * ''
Orculella astirakiensis ''Orculella astirakiensis'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as mollus ...
'' Gittenberger & Hausdorf 2004 * ''
Orculella brandti ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and ...
'' (Zilch 1960)Zilch, A. (1960). "Neue Landschnecken aus der Cyrenaica". ''Archiv für Molluskenkunde'' 89: 57-60. * ''
Orculella bulgarica bulgarica ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and ...
'' (P. Hesse 1915) * ''
Orculella bulgarica lamellata ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' Hausdorf, 1996Hausdorf, B. (1996). "Die Orculidae Asiens (Gastropoda: Stylommatophora)". ''Archiv für Molluskenkunde'' 125 (1/2): 1-86. * ''
Orculella caabensis ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' (Brandt 1956) * ''
Orculella connollyi ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and ...
'' (Brandt 1956) * ''
Orculella creantirudis ''Orculella creantirudis'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Orculidae. Geographic distribution ''O. creantirudis'' is endemic to Greece, where it occurs on the islands of Antikyth ...
'' Gittenberger & Hausdorf 2004 * ''
Orculella creticostata ''Orculella creticostata'' is a species of aerobic (air-breathing) land snail, a terrestrial pulmonate gastropod mollusc in the family Orculidae. Geographic distribution ''O. creticostata'' is endemic to Greece, where it occurs in the western p ...
'' Gittenberger & Hausdorf 2004 * '' Orculella cretilasithi'' Gittenberger & Hausdorf 2004 * ''
Orculella cretimaxima ''Orculella cretimaxima'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Orculidae Orculidae is a family of mostly minute, air-breathing, land snails; terrestrial pulmonate gastropod mollus ...
'' Gittenberger & Hausdorf 2004 * '' Orculella cretiminuta'' Gittenberger & Hausdorf 2004 * ''
Orculella cretioreina ''Orculella cretioreina'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Orculidae. Geographic distribution ''O. cretioreina'' is endemic to Greece, where it occurs at disjunct locations, usua ...
'' Gittenberger & Hausdorf 2004 * ''
Orculella critica ''Orculella critica'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Orculidae. Geographic distribution ''O. critica'' is native to Greece, where it is found in various areas of the mainland ...
'' (L. Pfeiffer 1856) * '' Orculella diensis'' Gittenberger & Hausdorf 2004 * ''
Orculella driana ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' (Klatenbach 1943) * ''
Orculella elongata ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' (Klatenbach 1943) * ''
Orculella exaggerata ''Orculella exaggerata'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Orculidae. Geographic distribution ''O. exaggerata'' is endemic to Greece, where it occurs on the islands of Kasos, ...
'' (Fuchs & Käufel 1936) * ''
Orculella farzughana ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' (Brandt 1956) * '' Orculella fodela'' Gittenberger & Hausdorf 2004 * '' Orculella franciscoi'' Gittenberger & Hausdorf 2004Gittenberger & Hausdorf (2004). "The Orculella species of the South Aegean island arc, a neglected radiation (Gastropoda, Pulmonata, Orculidae)". ''Basteria'' 68: 93-124. * ''
Orculella garzanensis ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' Schütt, 1996 * ''
Orculella gregoryi ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' (Brandt 1956) * '' Orculella? heterostropha commagenensis'' (Neubert, 1988) * '' Orculella? heterostropha heterostropha'' (O. Boettger, 1905) * '' Orculella ignorata'' Hausdorf, 1996 * ''
Orculella kaltenbachi ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' (Brandt 1956) * ''
Orculella mayi ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' (Brandt 1956) * ''
Orculella menkhorsti menkhorsti ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' Hausdorf, 1996 * '' Orculella menkhorsti sinistrorsa'' Hausdorf, 1996 * '' Orculella mesopotamica mesopotamica'' (Mousson, 1874) * '' Orculella mesopotamica riedeli'' Hausdorf, 1996 * ''
Orculella multidentata ''Orculella'' is a genus of air-breathing land snails, Terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of specie ...
'' (Klatenbach 1943) * ''
Orculella scalaris ''Orculella scalaris'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Orculidae. Geographic distribution ''O. scalaris'' is endemic to Greece, where it is restricted to the island of Dia, nort ...
'' Gittenberger & Hausdorf 2004 * ''
Orculella orientalis ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' (L. Pfeiffer 1861) * ''
Orculella pabsti ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' (Brandt 1956) * '' Orculella? palatalis'' (Pilsbry 1922) * ''
Orculella pfeiferi ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and ...
'' Hausdorf 1996 * ''
Orculella regimaensis ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the ...
'' (Brandt 1956) * ''
Orculella ruderalis urartaeica ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' Hausdorf 1996 * '' Orculella sirianocoriensis libanotica'' (Tristram 1865) * '' Orculella sirianocoriensis sirianocoriensis'' (Mousson 1854) * ''
Orculella striata ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' (Kaltenbach 1943) * ''
Orculella templorum templorum ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and ...
'' (Benoit 1862) * '' Orculella templorum tripolitana'' (Kaltenbach 1943) * ''
Orculella tingitana ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and ...
'' (Pallary 1918) * ''
Orculella tomlini ''Orculella'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Orculidae. The genus ''Orculella'' occurs in the whole Mediterranean region. The richest areas in terms of species are Turkey and the M ...
'' (Connolly 1942)Brandt, A. (1956). "Zur Orculidenfauna der Cyrenaica". ''Archiv für Molluskenkunde'' 85 1/3: 69-82.


References

Orculidae {{Orculidae-stub