The sheltopusik (''Pseudopus apodus''), also commonly called Pallas's glass lizard the European legless lizard, or the European glass lizard, is a species of large glass lizard found from Southern Europe to Central Asia. Pseudopus_apodus

Etymology

''Pseudopus apodus'' comes from Greek ψευδοποδος άποδος, literally meaning "fake-legged" or without legs. The common name "sheltopusik" comes from Russian желтопузик (''zheltopuzik''), which translates most directly as "yellow-bellied".

Taxonomy

The sheltopusik was previously included in the genus '' Ophisaurus'', but has since been placed in its own genus ''Pseudopus''. It was originally described in 1775 by Peter Simon Pallas as ''Lacerta apoda''. There are three subspecies: * ''P. a. apodus'' (Pallas, 1775) - the type subspecies, ranging from Crimea through Transcaucasia, east to Central Asia as far as Kazakhstan * ''P. a. levantinus'' Jablonski, Ribeiro-Junior, Meiri, Maza, Mikulíček & Janzik, 2021 - restricted to the Levant, from southern Turkey south to Israel * ''P. a. thracicus'' (Obst, 1978) - the westernmost subspecies, ranging from Croatia to Greece, east to western Turkey Despite only being described in 2021, ''P. a. levantinus'' is the largest and most genetically diverse of the subspecies, indicating an older and more complex evolutionary history compared to its sister subspecies.

Description

The sheltopusik can reach a length of . It is tan colored, paler on the ventral surface and the head, with a ring-like/segmented appearance that makes it look like a giant earthworm with a distinctive fold of skin down each side called a lateral groove. Small (2-mm) rear legs are sometimes visible near the

cloaca In animal anatomy, a cloaca ( ), plural cloacae ( or ), is the posterior orifice that serves as the only opening for the digestive, reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles and birds, a ...

. Though the legs are barely discernible, the sheltopusik can be quickly distinguished from a snake by its ears, eyelids, and ventral scales.

Habitat and behaviour

''P. apodus'' inhabits open country, such as short grassland or sparsely wooded hills. It consumes

arthropods Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

and small mammals. Snails and slugs appear to be its favorite prey, which may explain why it is particularly active in wet weather, although it prefers a dry habitat. Breaking through the shells of snails is an especially easy task due to their teeth and jaw structure.

Defensive behaviour

Due to its size, the sheltopusik tends to respond to harassment by hissing, biting, and musking. It is less likely to drop off its tail than some other species that display caudal autotomy. However, these occasional displays of caudal autotomy are responsible for the name " glass lizard" (or "glass snake"). The released tail may break into pieces, leading to the myth that the lizard can shatter like glass and reassemble itself later. In reality, if the tail is lost, it grows back slowly, but is shorter and darker. The replacement tail may grow back to full length after an extended period of time. Ophisaurus apodus2

In captivity

Sheltopusiks are frequently available in the exotic pet trade, though rarely captive-bred. They do not typically tolerate a large amount of handling, but they adapt to captivity well, feeding on crickets,

meal worm Mealworms are the larval form of the yellow mealworm beetle, ''Tenebrio molitor'', a species of darkling beetle. Like all holometabolism, holometabolic insects, they go through four life stages: egg (biology), egg, larva, pupa, and imago, adult ...

s, small

mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...

, eggs, snails, or pieces of meat. They are even known to accept these meals from a keeper's tweezers, or even from their hands once they become used to captivity. However, sheltopusiks do get excited around food and have surprisingly powerful jaws. They make hardy captives, capable of living up to 50 years.

Reproduction

About 10 weeks after mating, the female ''P. apodus'' lays about eight eggs, which she hides under bark or a stone, and often guards them. The young hatch after 45 to 55 days. They are typically about long and usually start to eat after four days.

Relationship with humans

Remains of the Levant subspecies (''P. a. levantinus'') are known from Natufian sites in Israel, suggesting that it was eaten by the local population at the time.

References

External links

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