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The neotropic cormorant or olivaceous cormorant (''Nannopterum brasilianum'') is a medium-sized cormorant found throughout the American
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referr ...
and subtropics, from the middle Rio Grande and the
Gulf A gulf is a large inlet from the ocean into the landmass, typically with a narrower opening than a bay, but that is not observable in all geographic areas so named. The term gulf was traditionally used for large highly-indented navigable bodies ...
and
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
n coasts of the United States south through
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
to southern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
, where it is called by the indigenous name of ''biguá''. It also breeds on the
Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to 88% of the ar ...
,
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
and
Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...
. It can be found both at coasts (including some mangrove areas) and on inland waters. There are at least two subspecies: ''N. b. mexicanum'' from
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the countr ...
northwards and ''N. b. brasilianum'' further south. In
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
the neotropic cormorant is used by the
Uru people The Uru or Uros ( ure, Qhas Qut suñi) are an indigenous people of Bolivia. They live on an approximate and still growing 120 self-fashioned floating islands in Lake Titicaca near Puno. They form three main groups: the Uru-Chipaya, Uru-Murat ...
for
fishing Fishing is the activity of trying to catch fish. Fish are often caught as wildlife from the natural environment, but may also be caught from stocked bodies of water such as ponds, canals, park wetlands and reservoirs. Fishing techniques inclu ...
.


Taxonomy

The neotropic cormorant was documented in 1658 by the Dutch naturalist
Willem Piso Willem Piso (in Dutch Willem Pies, in Latin Gulielmus Piso, also called Guilherme Piso in Portuguese) (1611 in Leiden – 28 November 1678 in Amsterdam) was a Dutch physician and naturalist who participated as an expedition doctor in D ...
after travels in
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. This formed the basis for the formal description and naming of the species by Johann Friedrich Gmelin in 1789. He placed it with the petrels in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Procellaria Procellaria is a genus of Southern Ocean long-winged seabirds related to prions, and within the order Procellariiformes. The black petrel (''Procellaria parkinsoni'') ranges in the Pacific Ocean, and as far north as Central America. The specta ...
'' and coined the binomial name ''Procellaria brasiliana''. Many authors preferred to use
Alexander von Humboldt Friedrich Wilhelm Heinrich Alexander von Humboldt (14 September 17696 May 1859) was a German polymath, geographer, naturalist, explorer, and proponent of Romantic philosophy and science. He was the younger brother of the Prussian minister, ...
's 1805 description of ''Pelecanus olivaceus'' because the identity of Piso's birds was considered uncertain. Later, ornithological authorities such as the American Ornithologists' Union began to use ''Phalacrocorax brasilianus'' after Ralph Browning argued that Piso's description and paintings do indeed refer to the neotropic cormorant. The neotropic cormorant was formerly placed in the genus '' Phalacrocorax''. A molecular phylogenetic study of the cormorants published in 2014 found that the neotropic cormorant, the
double-crested cormorant The double-crested cormorant (''Nannopterum auritum'') is a member of the cormorant family of water birds. It is found near rivers and lakes, and in coastal areas, and is widely distributed across North America, from the Aleutian Islands in Al ...
and the
flightless cormorant The flightless cormorant (''Nannopterum harrisi''), also known as the Galapagos cormorant, is a cormorant endemic to the Galapagos Islands, and an example of the highly unusual fauna there. It is unique in that it is the only known cormorant th ...
formed a clade that was
sister A sister is a woman or a girl who shares one or more parents with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a family, familial relationship, it is sometimes used endearingly to r ...
to the genus ''
Leucocarbo ''Leucocarbo'' is a genus of birds in the family Phalacrocoracidae with the members commonly known as blue-eyed shags. This is a group of closely related cormorant taxa. Many have a blue, purple or red ring around the eye (not a blue iris); othe ...
''. The three species were therefore moved the resurrected genus ''
Nannopterum ''Nannopterum'' is a genus of cormorant comprising three species. They are found throughout the Americas, hence the common name American cormorants. These species were formerly classified in the genus '' Phalacrocorax''. A molecular phylogenet ...
'' that had been introduced in 1899 by English ornithologist
Richard Bowdler Sharpe Richard Bowdler Sharpe (22 November 1847 – 25 December 1909) was an English zoologist and ornithologist who worked as curator of the bird collection at the British Museum of natural history. In the course of his career he published several mo ...
to accommodate the
flightless cormorant The flightless cormorant (''Nannopterum harrisi''), also known as the Galapagos cormorant, is a cormorant endemic to the Galapagos Islands, and an example of the highly unusual fauna there. It is unique in that it is the only known cormorant th ...
. The genus name ''Nannopterum'' combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
''nannos'' meaning "dwarf" with ''pteron'' meaning "wing". Two subspecies are recognised: * ''N. b. mexicanum'' ( Brandt, JF, 1837) – inland and coastal Great Plains (central USA) from South Dakota and Kansas to Gulf of Mexico, and Mexico to Nicaragua; Bahamas and Cuba * ''N. b. brasilianum'' (Gmelin, JF, 1789) – inland and coastal Costa Rica to
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla ...


Description

This bird is long with a wingspan. Adults males weigh from , adult females less. Birds of the southern populations tend to be bigger than the more northerly birds. It is small and slender, especially compared to the larger, heavier-looking
double-crested cormorant The double-crested cormorant (''Nannopterum auritum'') is a member of the cormorant family of water birds. It is found near rivers and lakes, and in coastal areas, and is widely distributed across North America, from the Aleutian Islands in Al ...
. It has a long tail and frequently holds its neck in an S-shape. Adult plumage is mainly black, with a yellow-brown throat patch. During breeding, white tufts appear on the sides of the head, there are scattered white
filoplume Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier ...
s on the side of the head and the neck, and the throat patch develops a white edge. The upper wings are somewhat grayer than the rest of the body. Juveniles are brownish in color.


Behaviour

Unlike other cormorants, this bird can often be seen perching on wires.


Food and feeding

Its diet consists mainly of small
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
, but will also eat
tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found ...
s,
frog A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" ''Triadobatrachus'' is ...
s, aquatic
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...
s (such as dragonfly
nymph A nymph ( grc, νύμφη, nýmphē, el, script=Latn, nímfi, label= Modern Greek; , ) in ancient Greek folklore is a minor female nature deity. Different from Greek goddesses, nymphs are generally regarded as personifications of nature, are ...
s), and shrimp. Information about its prey is sparse, but inland birds seem to feed on small, abundant fish in ponds and sheltered inlets, less than in length, with an individual weight of a gram or two, such as ''
Poecilia ''Poecilia'' is a genus of fishes in the family Poeciliidae of the order Cyprinodontiformes. These livebearers are native to fresh, brackish and salt water in the Americas, and some species in the genus are euryhaline. A few have adapted to livi ...
'' species especially the
sailfin molly The sailfin molly (''Poecilia latipinna'') is a species of fish of the genus '' Poecilia''. They inhabit fresh, brackish, salt, and coastal waters from North Carolina to Texas and the Yucatán Peninsula of Mexico. Taxonomy The sailfin molly was ...
''Poecilia latipinna''. This cormorant forages for food by diving underwater, propelling itself by its feet. Its dives are brief, between 5 and 15 seconds. It is also known to forage in groups, with several birds beating the water with their wings to drive fish forward into shallows.


Breeding

Neotropic cormorants are monogamous and breed in
colonies In modern parlance, a colony is a territory subject to a form of foreign rule. Though dominated by the foreign colonizers, colonies remain separate from the administration of the original country of the colonizers, the '' metropolitan state'' ...
. The
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
is a platform of sticks with a depression in the center circled with twigs and grass. It is built a few meters above the ground or water in bushes or trees. Up to five chalky, bluish-white
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
are laid. Most pairs lay three eggs, but the mean number hatched is less than two. The eggs soon become nest-stained. Both sexes incubate for about 25–30 days, and both parents feed the young until around the 11th week. By week 12, they are independent. One brood is raised per year. This bird is largely a permanent resident, with some birds occasionally wandering north in the warmer months. File:Phalacrocorax brasilianus -Argentina -two flying-8.jpg, Flying in Argentina File:Neotropic Cormorants1.jpg, New Mexico File:Phalacrocorax brasilianus-cooling.jpg,
Rodrigo de Freitas Lagoon Rodrigo de Freitas Lagoon (Portuguese: ''Lagoa Rodrigo de Freitas'') is a lagoon in the district of Lagoa in the Zona Sul (South Zone) area of Rio de Janeiro. The lagoon is connected to the Atlantic Ocean, allowing sea water to enter by a ca ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
File:Neotropic Cormorants in flights.jpg, Two cormorants in flight, in the
Pearl Islands The Pearl Islands (Spanish: Archipiélago de las Perlas or Islas de las Perlas) is a group of 200 or more islands and islets (many tiny and uninhabited) lying about off the Pacific coast of Panama in the Gulf of Panama. Islands The most nota ...
of Panama


References

* Johnsgaard, P. A. (1993), ''Cormorants, darters and pelicans of the world''. Washington DC: Smithsonian Institution Press. * Kaufman, Kenn; ''Lives of North American Birds''. Houghton Mifflin Company, New York, NY (1996). * World Wildlife Fund. 2010
''Petenes mangroves''
eds. Mark McGinley, C.Michael Hogan & C. Cleveland. Encyclopedia of Earth. National Council for Science and the Environment. Washington DC * Alsop, Fred J. III; ''Birds of Texas''. Smithsonian Handbooks: DK Publishing, Inc. (2002).


External links

* *
Neotropic Cormorant
Cornell University of Ornithology * {{Taxonbar, from=Q849173 Neotropic comorant Birds of the Americas Birds of the Rio Grande valleys Birds of Central America Birds of South America
Neotropic cormorant The neotropic cormorant or olivaceous cormorant (''Nannopterum brasilianum'') is a medium-sized cormorant found throughout the American tropics and subtropics, from the middle Rio Grande and the Gulf and Californian coasts of the United States so ...
Neotropic cormorant The neotropic cormorant or olivaceous cormorant (''Nannopterum brasilianum'') is a medium-sized cormorant found throughout the American tropics and subtropics, from the middle Rio Grande and the Gulf and Californian coasts of the United States so ...
Taxobox binomials not recognized by IUCN