''Offacolus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

euchelicerate The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mit ...

, a group of

chelicerate The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mite ...

arthropods Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

. Its only species, ''O. kingi'', has been found in deposits from the

Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...

period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

(

Homerian In the geologic timescale, the Homerian is an age of the Wenlock Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that is comprehended between 430.5 ± 0.7 Ma and 427.4 ± 0.5 Ma (million years ago), approximately. Th ...

epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...

) in the

Wenlock Series Wenlock may refer to: Places United Kingdom * Little Wenlock, a village in Shropshire * Much Wenlock, a town in Shropshire ** (Much) Wenlock (UK Parliament constituency) ** Wenlock Priory, a 7th/12th-century monastery * Wenlock Basin, a canal ba ...

Lagerstätte of Herefordshire,

England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...

. It is the only member of the

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Offacolidae, and classified as a basal ("primitive") genus in the

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

Euchelicerata, along with ''

Dibasterium ''Dibasterium'' is an extinct genus of euchelicerate, a group of chelicerate arthropods. Fossils of the single and type species, ''D. durgae'', have been discovered in the Coalbrookdale Formation of the Middle Silurian period (Homerian age) in ...

'' and

Prosomapoda Prosomapoda is a clade of euchelicerates including the groups Xiphosura (horseshoe crabs) and Planaterga (a group comprising bunodids, pseudoniscids, chasmataspidids, eurypterids and arachnids), as well as several basal synziphosurid genera ...

. The genus is named after

Offa Offa (died 29 July 796 AD) was King of Mercia, a kingdom of Anglo-Saxon England, from 757 until his death. The son of Thingfrith and a descendant of Eowa, Offa came to the throne after a period of civil war following the assassination of Æt ...

, a king from the ancient kingdom of

Mercia la, Merciorum regnum , conventional_long_name=Kingdom of Mercia , common_name=Mercia , status=Kingdom , status_text=Independent kingdom (527–879)Client state of Wessex () , life_span=527–918 , era=Heptarchy , event_start= , date_start= , ye ...

, and ''colus'', a person who dwelled among (this time referring to) the

Offa's Dyke Offa's Dyke ( cy, Clawdd Offa) is a large linear earthwork that roughly follows the border between England and Wales. The structure is named after Offa, the Anglo-Saxon king of Mercia from AD 757 until 796, who is traditionally believed to h ...

. The species name honors Robert Joseph King, a British

mineralogist Mineralogy is a subject of geology specializing in the scientific study of the chemistry, crystal structure, and physical (including optical) properties of minerals and mineralized artifacts. Specific studies within mineralogy include the proces ...

who found the fossils of ''Offacolus''. 20201122 Offacolus kingi ventral appendages

Similar to ''Dibasterium'', ''Offacolus'' possess limb-like exopods (outer limb branches) on appendage II to V, a character suggest to be

plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and ...

(observable in the putative stem-chelicerate taxon Habeliida) and lost within the prosomapod clade.

Classification

''Offacolus'' was originally described as an

arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

with

affinities, with detailed redescription done by Sutton ''et al.'' 2002 further suggested it to be unambiguously a chelicerate arthropod. ''Offacolus'' is the only

in the

Offacolidae, classified in the

Euchelicerata The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mite ...

together with the genus ''

'' and the clade

. A phylogenetic analysis (the results presented in a cladogram below) conducted by James Lamsdell in 2013 on the relationships within the Xiphosura and the relations to other closely related groups concluded that the Xiphosura, as presently understood, was

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

(a group sharing a

last common ancestor In biology and genetic genealogy, the most recent common ancestor (MRCA), also known as the last common ancestor (LCA) or concestor, of a set of organisms is the most recent individual from which all the organisms of the set are descended. The ...

but not including all descendants of this ancestor) and thus not a valid phylogenetic group. ''Offacolus'' was recovered as the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

(closest relative) of Prosomapoda.

References

{{reflist Euchelicerata Fossils of England Silurian arthropods of Europe Silurian animals Wenlock first appearances Homerian extinctions Fossil taxa described in 2000