''Evernia prunastri'', also known as oakmoss, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...

. Oakmoss grows primarily on the trunk and branches of

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

trees, but is also commonly found on the bark of other deciduous trees and

conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...

s such as fir and

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...

. The

thalli Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were ...

of oakmoss are short (3–4 cm in length) and bushy, and grow together on bark to form large clumps. Oakmoss thallus is flat and strap-like. They are also highly branched, resembling the form of

antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...

s. The colour of oakmoss ranges from green to a greenish-white when dry, and dark olive-green to yellow-green when wet. The texture of the thalli is rough when dry and rubbery when wet. It is used extensively in modern

perfumery Perfume (, ; french: parfum) is a mixture of fragrant essential oils or aroma compounds (fragrances), fixatives and solvents, usually in liquid form, used to give the human body, animals, food, objects, and living-spaces an agreeable scent. Th ...

Commercial uses

Oakmoss is commercially harvested in countries of South-Central Europe and usually exported to the

Grasse Grasse (; Provençal dialect, Provençal oc, Grassa in classical norm or in Mistralian norm ; traditional it, Grassa) is the only Subprefectures in France, subprefecture of the Alpes-Maritimes Departments of France, department in the Provence- ...

region of France where its fragrant compounds are extracted as oakmoss absolutes and extracts. These raw materials are often used as perfume fixatives and form the base notes of many fragrances. They are also key components of

Fougère ''Fougère'', , is one of the main olfactive families of perfumes. The name comes from the French language word for "fern". ''Fougère'' perfumes are made with a blend of fragrances: top-notes are sweet, with the scent of lavender flowers; ...

and

Chypre Chypre ( or ) is the name of a family (or ''concept'') of perfumes that are characterised by an accord composed of citrus top notes, a middle centered on cistus labdanum, and a mossy-animalic set of basenotes derived from oakmoss. Chypre perfumes ...

class perfumes. The lichen has a distinct and complex odor and can be described as woody, sharp and slightly sweet. Oakmoss growing on pines have a pronounced

turpentine Turpentine (which is also called spirit of turpentine, oil of turpentine, terebenthene, terebinthine and (colloquially) turps) is a fluid obtained by the distillation of resin harvested from living trees, mainly pines. Mainly used as a special ...

odor that is valued in certain perfume compositions. In parts of Central

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

, oakmoss has been used as for

biomonitoring In analytical chemistry, biomonitoring is the measurement of the body burden of toxic chemical compounds, elements, or their metabolites, in biological substances. Often, these measurements are done in blood and urine. Biomonitoring is performed ...

the deposition of

heavy metals upright=1.2, Crystals of osmium, a heavy metal nearly twice as dense as lead">lead.html" ;"title="osmium, a heavy metal nearly twice as dense as lead">osmium, a heavy metal nearly twice as dense as lead Heavy metals are generally defined as ...

urban Urban means "related to a city". In that sense, the term may refer to: * Urban area, geographical area distinct from rural areas * Urban culture, the culture of towns and cities Urban may also refer to: General * Urban (name), a list of people ...

rural In general, a rural area or a countryside is a geographic area that is located outside towns and cities. Typical rural areas have a low population density and small settlements. Agricultural areas and areas with forestry typically are describ ...

, and

industrial Industrial may refer to: Industry * Industrial archaeology, the study of the history of the industry * Industrial engineering, engineering dealing with the optimization of complex industrial processes or systems * Industrial city, a city dominate ...

sites. Studies of bioaccumulation for

zinc Zinc is a chemical element with the symbol Zn and atomic number 30. Zinc is a slightly brittle metal at room temperature and has a shiny-greyish appearance when oxidation is removed. It is the first element in group 12 (IIB) of the periodi ...

lead Lead is a chemical element with the symbol Pb (from the Latin ) and atomic number 82. It is a heavy metal that is denser than most common materials. Lead is soft and malleable, and also has a relatively low melting point. When freshly cu ...

chromium Chromium is a chemical element with the symbol Cr and atomic number 24. It is the first element in group 6. It is a steely-grey, lustrous, hard, and brittle transition metal. Chromium metal is valued for its high corrosion resistance and hardne ...

cadmium Cadmium is a chemical element with the symbol Cd and atomic number 48. This soft, silvery-white metal is chemically similar to the two other stable metals in group 12, zinc and mercury. Like zinc, it demonstrates oxidation state +2 in most of ...

, and

copper Copper is a chemical element with the symbol Cu (from la, cuprum) and atomic number 29. It is a soft, malleable, and ductile metal with very high thermal and electrical conductivity. A freshly exposed surface of pure copper has a pinkis ...

in lichen samples were performed five times at regular intervals between November 2000 and December 2001. As expected, the rural areas showed smaller impact of those five heavy metals when compared to urban and industrial areas.

Health and safety information

Oakmoss should be avoided by people with known skin sensitization issues. Its use in perfumes is now highly restricted by

International Fragrance Association The International Fragrance Association (IFRA) is the global representative body of the fragrance industry. It seeks to represent the collective interests of the industry and promote the safe use of fragrances through regulation. The Association wa ...

regulations, and many scents have been reformulated in recent years with other chemicals substituted for oakmoss.

Conservation status

''Evernia prunastri'' is listed as critically endangered (CR) in

Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its s ...

, where it is found in only one location.Náttúrufræðistofnun Íslands celandic Institute of Natural History(1996).
Válisti 1: Plöntur.
' (in Icelandic) Reykjavík: Náttúrufræðistofnun Íslands. As of April 2021, it has not been evaluated by the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

Gallery

Image:Evernia_prunastri-1.jpg, Close-up photo showing branches with soredia

References

Parmeliaceae Lichen species Perfume ingredients Taxa named by Carl Linnaeus Lichens described in 1753 Fungi of Europe Fungi of North America Fungi of Iceland {{Lecanorales-stub

Commercial uses

Health and safety information

Conservation status

Gallery

See also

References