The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the

Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...

Period million years ago ( Mya) to the beginning of the present

Quaternary The Quaternary ( ) is the current and most recent of the three periods of the Cenozoic Era in the geologic time scale of the International Commission on Stratigraphy (ICS). It follows the Neogene Period and spans from 2.58 million years ...

Period Mya. The Neogene is sub-divided into two

epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...

s, the earlier

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

and the later

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Quaternary The Quaternary ( ) is the current and most recent of the three periods of the Cenozoic Era in the geologic time scale of the International Commission on Stratigraphy (ICS). It follows the Neogene Period and spans from 2.58 million years ...

. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). During this period,

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s and

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

s continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans (''

Homo habilis ''Homo habilis'' ("handy man") is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.31 million years ago to 1.65 million years ago (mya). Upon species description in 1964, ''H. habilis'' was highly ...

'') appeared in Africa near the end of the period. Some continental movements took place, the most significant event being the connection of

North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...

and

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

at the Isthmus of Panama, late in the Pliocene. This cut off the warm ocean currents from the Pacific to the Atlantic Ocean, leaving only the

Gulf Stream The Gulf Stream, together with its northern extension the North Atlantic Current, North Atlantic Drift, is a warm and swift Atlantic Ocean, Atlantic ocean current that originates in the Gulf of Mexico and flows through the Straits of Florida a ...

to transfer heat to the

Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five major oceans. It spans an area of approximately and is known as the coldest of all the oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, a ...

. The global climate cooled considerably throughout the Neogene, culminating in a series of continental

glaciations A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate betwe ...

in the

Period that follows.

Divisions

In ICS terminology, from upper (later, more recent) to lower (earlier): The

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Piacenzian The Piacenzian is in the international geologic time scale the upper stage (stratigraphy), stage or latest age (geology), age of the Pliocene. It spans the time between 3.6 ± 0.005 year#SI prefix multipliers, Ma and 2.588 ± 0.005 Ma (million yea ...

Age, preceded by *

Zanclean The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, and ...

Age The

Epoch is subdivided into six ages: * Messinian Age, preceded by *

Tortonian The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Torton ...

Age *

Serravallian The Serravallian is, in the geologic timescale, an age or a stage in the middle Miocene Epoch/Series, which spans the time between 13.82 Ma and 11.63 Ma (million years ago). The Serravallian follows the Langhian and is followed by the Tortonian ...

Age *

Langhian The Langhian is, in the International Commission on Stratigraphy, ICS geologic timescale, an age (geology), age or stage (stratigraphy), stage in the middle Miocene epoch (geology), Epoch/series (stratigraphy), Series. It spans the time between 15 ...

Age *

Burdigalian The Burdigalian is, in the geologic timescale, an age (geology), age or stage (stratigraphy), stage in the early Miocene. It spans the time between 20.43 ± 0.05 annum, Ma and 15.97 ± 0.05 Ma (million years ago). Preceded by the Aquitanian (sta ...

Age * Aquitanian Age In different geophysical regions of the world, other regional names are also used for the same or overlapping ages and other timeline subdivisions. The terms ''Neogene System'' (formal) and ''Upper Tertiary System'' (informal) describe the rocks deposited during the ''Neogene Period''.

Geography

The continents in the Neogene were very close to their current positions. The Isthmus of Panama formed, connecting

and

. The

Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...

continued to collide with

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

, forming the

Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...

. Sea levels fell, creating

land bridge In biogeography, a land bridge is an isthmus or wider land connection between otherwise separate areas, over which animals and plants are able to cross and Colonisation (biology), colonize new lands. A land bridge can be created by marine regre ...

s between

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

and

Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...

and between Eurasia and North America.

Climate

The global climate became seasonal and continued an overall drying and cooling trend which began at the start of the

. During the

Early Miocene The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene Epoch made up of two stages: the Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 Ma to 15.97 ± 0.05 Ma (million years ago). It was prece ...

, mid-latitude seawater and continental thermal gradients were already very similar to those of the present. The ice caps on both poles began to grow and thicken, and by the end of the period the first of a series of glaciations of the

current Ice Age The Late Cenozoic Ice Age,National Academy of Sciences - The National Academies Press - Continental Glaciation through Geologic Times https://www.nap.edu/read/11798/chapter/8#80 or Antarctic Glaciation began 33.9 million years ago at the Eocen ...

began.

Flora and fauna

Marine and continental flora and fauna have a modern appearance. The reptile group Choristodera went extinct in the early part of the period, while the amphibians known as

Allocaudata The Albanerpetontidae are an extinct family of small amphibians, native to the Northern Hemisphere during the Mesozoic and Cenozoic. The only members of the order Allocaudata, they are thought to be allied with living amphibians belonging to Lissa ...

disappeared at the end of it. Neogene also marked the end of the reptilian genera

Langstonia ''Langstonia'' (meaning " rocodileof Langston", in honor of paleontologist Wann Langston, Jr.) is an extinct genus of notosuchian crocodylomorph of the family Sebecidae. It lived in the middle Miocene (specifically in the Laventan land-mammal ...

and

Barinasuchus ''Barinasuchus'' (meaning " Barinas crocodile," in reference to where the type material was found) is an extinct genus of sebecid mesoeucrocodylian. Its fossils have been found in middle Eocene-age rocks of the Divisadero Largo Formation of Arg ...

, terrestrial predators that were the last surviving members of

Sebecosuchia Sebecosuchia is an extinct group of mesoeucrocodylian crocodyliforms that includes the families Sebecidae and Baurusuchidae. The group was long thought to have first appeared in the Late Cretaceous with the baurusuchids and become extinct in th ...

, a group related to crocodiles. The oceans were dominated by large carnivores like

megalodon Megalodon (''Otodus megalodon''), meaning "big tooth", is an extinct species of mackerel shark that lived approximately 23 to 3.6 million years ago (Mya), from the Early Miocene to the Pliocene epochs. It was formerly thought to be a membe ...

s and

livyatan ''Livyatan'' is an extinct genus of macroraptorial sperm whale containing one known species: ''L. melvillei''. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous ...

s, and 19 million years ago about 70% of all pelagic shark species disappeared.

Mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s and

s continued to be the dominant terrestrial vertebrates, and took many forms as they adapted to various habitats. The first

hominin The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The ...

s, the ancestors of humans may have appeared in southern Europe and migrated into Africa. The first humans (belonging to the species ''

'') appeared in Africa near the end of the period. About 20 million years ago

gymnosperms The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνό� ...

in the form of some

conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...

and

cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...

groups started to diversify and produce more species due to the changing conditions. In response to the cooler, seasonal climate, tropical plant species gave way to

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...

ones and grasslands replaced many forests. Grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as

horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...

antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...

, and

bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'', found only in North Ame ...

. Ice age mammals like the

mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks and, ...

s and

woolly rhinoceros The woolly rhinoceros (''Coelodonta antiquitatis'') is an extinct species of rhinoceros that was common throughout Europe and Asia during the Pleistocene epoch and survived until the end of the last glacial period. The woolly rhinoceros was a me ...

were common in

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58 plants expanded and reached ecological dominance in grasslands during the last 10 million years. Also

Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...

(daisies) went through a significant

adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...

Eucalyptus ''Eucalyptus'' () is a genus of over seven hundred species of flowering trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe Eucalypteae, including '' Corymbia'', they are commonly known as euca ...

fossil leaves occur in the

of New Zealand, where the genus is not native today, but have been introduced from Australia.

Disagreements

The Neogene traditionally ended at the end of the Pliocene Epoch, just before the older definition of the beginning of the

Period; many time scales show this division. However, there was a movement amongst geologists (particularly

marine geologist Marine geology or geological oceanography is the study of the history and structure of the ocean floor. It involves geophysical, geochemical, sedimentological and paleontological investigations of the ocean floor and coastal zone. Marine geology ...

s) to also include ongoing geological time (Quaternary) in the Neogene, while others (particularly terrestrial geologists) insist the Quaternary to be a separate period of distinctly different record. The somewhat confusing terminology and disagreement amongst geologists on where to draw what hierarchical boundaries is due to the comparatively fine divisibility of time units as time approaches the present, and due to geological preservation that causes the youngest sedimentary geological record to be preserved over a much larger area and to reflect many more environments than the older geological record. By dividing the

Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...

Era into three (arguably two) periods (

, Neogene,

) instead of seven epochs, the periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic Eras. The

International Commission on Stratigraphy The International Commission on Stratigraphy (ICS), sometimes referred to unofficially as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigra ...

(ICS) once proposed that the Quaternary be considered a sub-era (sub-erathem) of the Neogene, with a beginning date of 2.58 Ma, namely the start of the Gelasian Stage. In the 2004 proposal of the ICS, the Neogene would have consisted of the

and

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58International Union for Quaternary Research The International Union for Quaternary Research (INQUA) was founded in 1928. It has members from a number of scientific disciplines who study the environmental changes that occurred during the glacial ages, the last 2.6 million years. One go ...

(INQUA) counterproposed that the Neogene and the Pliocene end at 2.58 Ma, that the Gelasian be transferred to the Pleistocene, and the Quaternary be recognized as the third period in the Cenozoic, citing key changes in Earth's climate, oceans, and biota that occurred 2.58 Ma and its correspondence to the Gauss-Matuyama magnetostratigraphic boundary.Clague, John ''et al.'' (2006) "Open Letter by INQUA Executive Committee"
''Quaternary Perspective, the INQUA Newsletter'' International Union for Quaternary Research 16(1) In 2006 ICS and INQUA reached a compromise that made Quaternary a subera, subdividing Cenozoic into the old classical Tertiary and Quaternary, a compromise that was rejected by International Union of Geological Sciences because it split both Neogene and Pliocene in two. Following formal discussions at the 2008 International Geological Congress in Oslo, Norway, the ICS decided in May 2009 to make the Quaternary the youngest period of the Cenozoic Era with its base at 2.58 Mya and including the Gelasian Age, which was formerly considered part of the Neogene Period and Pliocene Epoch. Thus the Neogene Period ends bounding the succeeding Quaternary Period at 2.58 Mya.

References

External links

* {{Authority control Geological periods