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The Namaqua sandgrouse (''Pterocles namaqua''), is a species of ground-dwelling bird in the
sandgrouse Sandgrouse is the common name for Pteroclidae , a family of sixteen species of bird, members of the order Pterocliformes . They are traditionally placed in two genera. The two central Asian species are classified as ''Syrrhaptes'' and the other ...
family. It is found in arid regions of south-western Africa.


Taxonomy

The Namaqua sandgrouse was formally described in 1789 by the German naturalist
Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich Gmelin Ferdinand Christoph Oetinger , academic_advisors = , docto ...
in his revised and expanded edition of Carl Linnaeus's ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nom ...
''. He placed it with all the grouse like birds in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
''
Tetrao ''Tetrao'' is a genus of birds in the grouse subfamily known as capercaillies. They are some of the largest living grouse. Taxonomy The genus ''Tetrao'' was introduced in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Tetrao namaqua''. Gmelin based his description on the "Namaqua grous" that had been described in 1783 by the English ornithologist John Latham in his ''A General Synopsis of Birds''. The Namaqua sandgrouse is now placed with 13 other species in genus '' Pterocles'' that was introduced in 1815 by the Dutch zoologist
Coenraad Jacob Temminck Coenraad Jacob Temminck (; 31 March 1778 – 30 January 1858) was a Dutch aristocrat, zoologist and museum director. Biography Coenraad Jacob Temminck was born on 31 March 1778 in Amsterdam in the Dutch Republic. From his father, Jacob Temmin ...
. The genus name combines the Ancient Greek ''pteron'' meaning "wing" with ''-klēs'' meaning "notable" or "splendid". The specific epithet ''namaqua'' is from Namaqualand, now
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
, the type locality. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
: no
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised.


Description

The sandgrouse is a medium-sized bird with a plump body, small head and short legs. It grows to a length of about . The male has an orangish buff head, throat and chest delineated by a conspicuous narrow band of white and dark brown. The back and wings are mottled brown with large white specks and there are two long black filaments extending from the olive-brown tail. The colouring of the female and juvenile is more cryptic being generally various shades of brown patterned with white specks. It could be confused with the
double-banded sandgrouse The double-banded sandgrouse (''Pterocles bicinctus'') is a species of ground-living bird in the family Pteroclidae. It is found in arid parts of southern Africa. Subspecies There are three subspecies: ''P. b. ansorgei'' is found in south west ...
(''Pterocles bicinctus'') and
Burchell's sandgrouse Burchell's sandgrouse (''Pterocles burchelli'') is a species of bird in the family Pteroclidae. It is found in arid and semi-arid regions of southern Africa. The name of this bird commemorates the English naturalist William John Burchell. Descr ...
(''Pterocles burchelli''), which share the same range.


Distribution and habitat

The Namaqua sandgrouse can be found in various arid parts of South Africa and its neighbouring lands. It is common in the
Kalahari Desert The Kalahari Desert is a large semi-arid sandy savanna in Southern Africa extending for , covering much of Botswana, and parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African Namib coastal ...
, the
Nama Karoo Nama Karoo is a xeric shrubland ecoregion located on the central plateau of South Africa and Namibia. It occupies most of the interior of the western half of South Africa and extends into the southern interior of Namibia. Climate The climate te ...
(in central and western South Africa), and in parts of the
Western Cape The Western Cape is a province of South Africa, situated on the south-western coast of the country. It is the fourth largest of the nine provinces with an area of , and the third most populous, with an estimated 7 million inhabitants in 2020 ...
. In addition, it is also found in
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
(especially in the
Namib Desert The Namib ( ; pt, Namibe) is a coastal desert in Southern Africa. The name is of Khoekhoegowab origin and means "vast place". According to the broadest definition, the Namib stretches for more than along the Atlantic coasts of Angola, Nami ...
), Zimbabwe, Botswana and Angola. It favours deserts and other arid areas. In order to survive, this bird needs only seeds, some gravel, and easy access to some sort of fresh water source. Their habitat usually has rough vegetation and tussock-type grasses.


Behaviour

Outside the breeding season, the sandgrouse are gregarious. The birds converge on watering holes in the early morning and several dozens or even hundreds of individuals may congregate in one place. They also tend to spend the night in groups, congregating about an hour before dusk. They split up during the day into much smaller groups to feed.


Feeding

Their principal diet is seeds but they also eat leaves, flowers, small fruit, insects and molluscs. They forage by exploring loose soil with their beaks and flicking it away sideways.


Breeding

Breeding takes place at any time of the year and is dependent on rainfall. Usually the nests are solitary but sometimes several pairs of birds choose sites near each other. The nest is a scrape in the earth, scantily lined with dried plant material. Two or three pinkish-grey eggs with brown markings are laid over the course of a few days. Incubation starts after the last egg has been laid and lasts about 22 days. The female does the incubation by day and the male does a longer shift at night, starting about two hours before sunset and finishing two hours after dawn. The chicks are
precocial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
and able to leave the nest on the day they are hatched. The male brings them water absorbed on the specially adapted feathers of his breast. The chicks grow rapidly; they are fully feathered at three weeks and able to fly at six.


Status and threats

The species is common within its range and is considered to be of
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
by the IUCN Red List of Threatened Species. The birds are at risk of
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
by
mongoose A mongoose is a small terrestrial carnivorous mammal belonging to the family Herpestidae. This family is currently split into two subfamilies, the Herpestinae and the Mungotinae. The Herpestinae comprises 23 living species that are native to s ...
s while they are young. Sheep farmers kill
birds of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
and
jackal Jackals are medium-sized canids native to Africa and Eurasia. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed ...
s to protect their flocks and this may have resulted in an increase in the mongoose population and consequently a diminution in the number of sandgrouse chicks that survive. Other predators that prey on the Namaqua sandgrouse include the
booted eagle The booted eagle (''Hieraaetus pennatus'', also classified as ''Aquila pennata'') is a medium-sized mostly migratory bird of prey with a wide distribution in the Palearctic and southern Asia, wintering in the tropics of Africa and Asia, with a ...
(''Aquila pennatus''), the
tawny eagle The tawny eagle (''Aquila rapax'') is a large bird of prey. Like all eagles, it belongs to the family ''Accipitridae''. Its heavily feathered legs mark it as a member of the subfamily Aquilinae, also known as booted eagles.Helbig, A. J., Kocum, ...
(''Aquila rapax''), the
lanner falcon The lanner falcon (''Falco biarmicus'') is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A ...
(''Falco biarmicus'') and the
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey ( raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey ba ...
(''Falco peregrinus''). The Namaqua sandgrouse is a host of the
Acanthocephalan Acanthocephala (Greek , ', thorn + , ', head) is a phylum of parasitic worms known as acanthocephalans, thorny-headed worms, or spiny-headed worms, characterized by the presence of an eversible proboscis, armed with spines, which it uses to ...
intestinal parasite ''
Moniliformis kalahariensis Moniliformidae is a family of parasitic spiny-headed (or thorny-headed) worms. It is the only family in the Moniliformida order and contains three genera: ''Australiformis'' containing a single species, ''Moniliformis'' containing eighteen speci ...
''.


References


External links

* Namaqua sandgrouse �
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q127964 Pterocles Birds of Southern Africa Birds described in 1789 Taxa named by Johann Friedrich Gmelin Taxonomy articles created by Polbot