Nyctophilus Arnhemensis
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''Nyctophilus arnhemensis'', known as the northern or Arnhem long-eared bat, is a species of
Chiroptera Bats are mammals of the Order (biology), order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in fli ...
(bats) native to northern regions of Australia. The distribution range is from north-western Queensland to northern Western Australia.


Taxonomy

The description of the species was first published in 1959 by David H. Johnson, the result of examination of mammal specimens the author collected on a 1948 scientific expedition backed by American and Australia institutions. The type specimen was collected near
Yirrkala Yirrkala is a small community in East Arnhem Region, Northern Territory, Australia, southeast of the large mining town of Nhulunbuy, Northern Territory, Nhulunbuy, on the Gove Peninsula in Arnhem Land. Its population comprises predominantly ...
, in the
Arnhem Land Arnhem Land is a historical region of the Northern Territory of Australia, with the term still in use. It is located in the north-eastern corner of the territory and is around from the territory capital, Darwin. In 1623, Dutch East India Compan ...
region, and placed at the
US National Museum The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7 ...
. The taxon is widely recognised. The epithet ''arnhemensis'' indicates the origin of Johnson's specimen. Common names regionally distinguish this species of ''Nyctophilus'': the northern, Arnhem, or Arnhem Land, 'long-eared bat', or as Arnhem nyctophilus. The species is not restricted in range to the Arnhem Land, and occurs beyond this region of northern Australia. The vernacular northern long-eared bat is also used in reference to '' Nyctophilus daedalus''.


Description

A species of ''
Nyctophilus ''Nyctophilus'' is a genus of the vespertilionids or vesper bats. They are often termed Australian big-eared bats or long-eared bats, as the length of their ears often greatly exceeds that of the head. This genus occurs in the New Guinean-Austr ...
'' of intermediate size. The measurement of the tibia is 36 to 40 millimetres, and weight range is 5 to 8 grams. The hair of the back is a mid brown, rusty colour, ventral fur is lighter. An indistinct ridge of flesh, located behind the snout, is shallowly incised. The litter size is thought to be up to two young. The shorter wing arrangement is more broad, allowing it to manoeuvre more slowly through densely vegetated environments that are unavailable to micro-bats of the region. The hunting and foraging technique is to patrol branches and foliage seeking insects.


Habitat and distribution

''Nyctophilus arnhemensis'' is found inhabiting
mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
, woodland and forest, and favours roosts in thick vegetation, beneath loose cover near a tree trunk. They reside under the papery bark of
melaleuca ''Melaleuca'' () is a genus of nearly 300 species of plants in the myrtle family, Myrtaceae, commonly known as paperbarks, honey-myrtles or tea-trees (although the last name is also applied to species of '' Leptospermum''). They range in size ...
species and especially favour
pandanus ''Pandanus'' is a genus of monocots with some 750 accepted species. They are palm-like, dioecious trees and shrubs native to the Old World tropics and subtropics. The greatest number of species are found in Madagascar and Malaysia. Common names ...
in
riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...
zones. It is locally common, but limited by the amount of suitable habitat; they have been reported occupying residential roof structures. There is a preference for mangrove, especially west of the
Dampier Peninsula The Dampier Peninsula is a peninsula located north of Broome and Roebuck Bay in Western Australia. It is surrounded by the Indian Ocean to the west and north, and King Sound to the east. It is named after the mariner and explorer William Damp ...
. Freshwater sites include lagoons and waterholes. The distribution range is at tropical regions across the north of the continent, near fresh or saline waters, at coastal areas and offshore island. This range extends beyond the Kimberley region to the west and to the
Gulf of Carpentaria The Gulf of Carpentaria (, ) is a large, shallow sea enclosed on three sides by northern Australia and bounded on the north by the eastern Arafura Sea (the body of water that lies between Australia and New Guinea). The northern boundary is ...
in the east, an outlying record at Cape York (the tip of
Cape York Peninsula Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupación ...
) was tentatively identified as this species. However, as with many vespertilionid bats of northern Australia, the population seem to have become geographically isolated by the Gulf of Carpentaria, which lacks caves and suitable trees for roosts. They occur at islands of the north, Melville Island, the
Bonaparte Archipelago The Bonaparte Archipelago is a group of islands off the coast of Western Australia in the Kimberley region, within the Shire of Wyndham-East Kimberley. The closest inhabited place is Kalumburu located about to the east of the island group. Th ...
,
Groote Eylandt Groote Eylandt ( Anindilyakwa: ''Ayangkidarrba'' meaning "island" ) is the largest island in the Gulf of Carpentaria and the fourth largest island in Australia. It was named by the explorer Abel Tasman in 1644 and is Dutch for "Large Island" in ...
, and the
Sir Edward Pellew Group The Sir Edward Pellew Group of Islands is situated in the south-west corner of the Gulf of Carpentaria, off the coast of the Northern Territory, Australia. History They were named in 1802 by Matthew Flinders in honour of Sir Edward Pellew, ...
. The species has fared better in pandanus environs, and remains common where the once populous hoary wattled ''
Chalinolobus nigrogriseus The hoary wattled bat (''Chalinolobus nigrogriseus'') is a species of vesper bat found in northern Australia and Papua New Guinea. Two subspecies are currently recognised: * ''C. n. nigrogriseus'' ( Gould, 1852) * ''C. n. rogersi'' (Thomas Tho ...
'' seemed vulnerable to the more frequent burns; the ability to detect and evade advancing fire-fronts is assumed to have given the species an advantage. ''N. arnhemensis'' remains well hidden beneath dead fronds of the pandanus and frequently relocates, for reason of hygiene or evading a potential predator. The wetland habitat of pandanus provides large and diverse sources of food for this bat. The species has a large population that is presumed level or increasing, and identified as 'least concern' on the international
red list The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
. The bats are vulnerable to the consequences of altered land use, to agricultural and pastoralist activities, primarily the removal of their roosting and foraging habitat.


References

{{Taxonbar, from=Q1830467 Bats of Australia Mammals of Western Australia Mammals of the Northern Territory Mammals of Queensland Nyctophilus Mammals described in 1959