Noturus Flavus
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''Noturus flavus'', the stonecat, is a North American freshwater
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Ictaluridae. The common name is due to its habit of hiding near or under stones in fast-moving water.


Anatomy

Stonecats are small, slender, flat-headed catfishes, with the adipose fin keel-like and continuous with the caudal fin except for a shallow notch. While the dorsal and pectoral spines of most members of this family cause wounds that irritate, the stonecat has venom glands, particularly on the bases of the pectoral spines, that cause extreme pain similar to that of a wasp sting. The pectoral fin lacks any posterior serrae. Anal fin rays number 15 to 18, pectoral fin rays 9 to 11, and pelvic fin rays 8 to 10. The caudal fin rays number 55 to 67. The upper jaw projects beyond the lower jaw and the tooth pad on the upper jaw has a narrow, crescent-shaped extension on each side. The premaxillary band of teeth has lateral backward extensions. The
skin Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other cuticle, animal coverings, such as the arthropod exoskeleton, have diffe ...
of the stonecat is thick and is yellowish-brown in color. The sides of the head shade to yellow. The belly is whitish. The stonecat has two forms. In the Cumberland drainage in
Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...
, a scientifically undescribed form possesses two light bars (perpendicular to body length) on its nape. In other areas, a patch exists in place of the bars. In both forms, the stonecat has a white spot at the rear of the dorsal fin base and one on the upper edge of the caudal fin. The rear of the pectoral spine has either no or a few weak teeth.Noturus flavus


Size/age

Stonecats typically reach four to eight inches in length, but can reach 12 inches, at weights of 0.22 to 1.1 lbs. Typically, Stonecats live five to six years.


Distribution

The stonecat has a widespread distribution. Stonecats exist in the
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lakes ...
, the
St. Lawrence River The St. Lawrence River (french: Fleuve Saint-Laurent, ) is a large river in the middle latitudes of North America. Its headwaters begin flowing from Lake Ontario in a (roughly) northeasterly direction, into the Gulf of St. Lawrence, connecting ...
, drainages of
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
, and the Mississippi River basin. Stonecats can also be found from the
Hudson River The Hudson River is a river that flows from north to south primarily through eastern New York. It originates in the Adirondack Mountains of Upstate New York and flows southward through the Hudson Valley to the New York Harbor between N ...
drainage of New York west to the Red River drainage of Hudson Bay. Stonecats inhabit the drainage of the
Mississippi River The Mississippi River is the second-longest river and chief river of the second-largest drainage system in North America, second only to the Hudson Bay drainage system. From its traditional source of Lake Itasca in northern Minnesota, it f ...
basin from Quebec to Alberta, southerly to northern Alabama and Mississippi and westerly to northeastern Oklahoma. In Colorado, stonecats are present in
St. Vrain Creek St. Vrain Creek (often known locally as the St. Vrain River) is a tributary of the South Platte River, approximately long,U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map accessed March 25, 2011 ...
near
Longmont, Colorado The City of Longmont is a home rule municipality located in Boulder and Weld counties, Colorado, United States. Longmont is located northeast of the county seat of Boulder and north-northwest of the Colorado State Capitol in Denver. Longmont ...
and in the
Republican River The Republican River is a river in the central Great Plains of North America, rising in the High Plains (United States), High Plains of eastern Colorado and flowing east U.S. Geological Survey. National Hydrography Dataset high-resolution flowline ...
south of Wray, Colorado."Critical Thermal Maxima of Adult Stonecats", pages 8 to 11, Volume 30, Issue 3, May/June 2013, ''Colorado Water'' published by Colorado State University


Ecology


Habitat

Stonecats live in
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
environments, and are found from large creeks to small rivers. They occasionally are found in tiny creeks, or rivers as large as the Lower Mississippi. Stonecats occupy gently to fast-moving riffle areas with rocky substrates. They spend the majority of their time in moderate-moving, shallow riffles, but can also be found in deeper water (2 to 3 meters deep). Stonecat inhabit natural lakes, such as
Lake Erie Lake Erie ( "eerie") is the fourth largest lake by surface area of the five Great Lakes in North America and the eleventh-largest globally. It is the southernmost, shallowest, and smallest by volume of the Great Lakes and therefore also has t ...
, where it prefers rock and gravel bars that are subject to significant wave action.


Diet

The stonecat is a Benthic zone, benthic, opportunistic feeder, using its sensitive barbels during the night to search for food on the river bottom. Stonecats eat a diversity of food items, such as aquatic insect larvae (e.g., mayflies), mollusks, minnows, fish eggs, isopods, amphipods, crayfish, plant material, worms and chilopods.


Reproduction

Females mature in three to four years and a mean standard length of 4.7 inches. Stonecats form monogamous pairs for breeding, and spawn when water temperatures reach 25 °C. Clutches are guarded by males under large, flat rocks in pools or crests of riffles. Rocks used as spawning cover averaged 200 square inches and were found in water depths averaging 34 inches. The eggs are amber-yellow and are very large, ranging between 3.5 and 4 mm in diameter, with the whole egg mass enveloped by a gelatinous material. A female stonecat may produce between 200 and 1,200 eggs per year. Stonecats exhibit parental care, with the male or both sexes guarding the clutch, until the young head to shallower, calmer streams and waters to mature.


Etymology

The genus name ''Noturus'', meaning "back tail", refers to the fusion of the adipose and caudal fins. The specific epithet ''flavus'' meaning "yellow", refers to the color distinction.


Importance/conservation status

Stonecats serve as indicators of
water quality Water quality refers to the chemical, physical, and biological characteristics of water based on the standards of its usage. It is most frequently used by reference to a set of standards against which compliance, generally achieved through tr ...
: they are not present in highly polluted or heavily silted areas. Stonecats are a valuable
indicator species A bioindicator is any species (an indicator species) or group of species whose function, population, or status can reveal the qualitative status of the environment. The most common indicator species are animals. For example, copepods and other sma ...
to humans, as they may also be useful as a marker for water temperature. The US Endangered Species Act lists the status of ''N. flavus'' as not threatened or no special status, meaning that this species faces no risk of extinction.


References


Bibliography

*Eddy, Samuel and Surber Thaddeus. ''Northern Fishes'' The University of Minnesota Press, Minneapolis, 1943 pp. 160–161, ASIN: B001OJ4F5W. *Phillips, L. Gary, Schmid, D. William, and Underhill, C. James. ''Fishes of the Minnesota Region'' The University of Minnesota Press, Minneapolis, 1982 pp. 179–180, . *Page, M. Lawrence and Burr, M. Brooks. ''Freshwater Fishes'' Houghton Mifflin Company, Boston, New York, 1999 pp. 199–200, Plate 26, . *


External links


Stonecat pictures
* ttp://fieldguide.mt.gov/detail_AFCKA02070.aspx StoneCat facts {{Taxonbar, from=Q3752427 Noturus Fish of the Great Lakes Freshwater fish of the United States Fauna of the Plains-Midwest (United States) Fish described in 1818 Taxa named by Constantine Samuel Rafinesque