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The Niger Delta red colobus (''Piliocolobus epieni'') is a critically endangered species of
colobus monkey Black-and-white colobuses (or colobi) are Old World monkeys of the genus ''Colobus'', native to Africa. They are closely related to the red colobus monkeys of genus '' Piliocolobus''. There are five species of this monkey, and at least eight subs ...
endemic to the western part of the
Niger Delta The Niger Delta is the delta of the Niger River sitting directly on the Gulf of Guinea on the Atlantic Ocean in Nigeria. It is located within nine coastal southern Nigerian states, which include: all six states from the South South geopolitical ...
. It is threatened by hunting and
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
.


Taxonomy

From the time it first became known to science (in 1993) until 2007 or 2008, it was considered a subspecies of the
western red colobus The western red colobus (''Piliocolobus badius''), also known as the bay red colobus, rust red colobus or Upper Guinea red colobus, is a species of Old World monkey in West African forests from Senegal to Ghana. All other species of red colobuses ...
(''Procolobus badius'') and more recently
Pennant's colobus Pennant's colobus or Pennant's red colobus (''Piliocolobus pennantii'') is a species of tree-dwelling primate in the family Cercopithecidae. It is endemic to tropical Central Africa. Three subspecies have traditionally been recognised but its dis ...
(''Procolobus pennantii''), and its trinomial name was ''Procolobus badius epieni'' or ''Procolobus pennantii epieni''.
Colin Groves Colin Peter Groves (24 June 1942 – 30 November 2017) was a British-Australian biologist and anthropologist. Groves was Professor of Biological Anthropology at the Australian National University in Canberra, Australia. Education Born in Englan ...
recognized the Niger Delta red colobus as a full species in 2007, although Groves considered all the red colobus monkeys, including the Niger Delta red colobus to be in the genus '' Piliocolobus'', rather than '' Procolobus'' (this nomenclature has been followed here). However, other authors consider ''Piliocolobus'' to be a
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of ''Procolobus''.


Description

The Niger Delta red colobus is black on top from the head to the rump, becoming orange-brown on the sides and outer legs. The undersides and inner legs, and most of the arms are white. The hands and feet are black. The tail is red-brown on top and chestnut or maroon below, becoming darker towards the tip. It has white whiskers.


Distribution

The Niger Delta red colobus is found only in the western part of the
Niger Delta The Niger Delta is the delta of the Niger River sitting directly on the Gulf of Guinea on the Atlantic Ocean in Nigeria. It is located within nine coastal southern Nigerian states, which include: all six states from the South South geopolitical ...
. It is restricted to marsh forest, that is, forest with a high
water table The water table is the upper surface of the zone of saturation. The zone of saturation is where the pores and fractures of the ground are saturated with water. It can also be simply explained as the depth below which the ground is saturated. T ...
year round but no significant flooding. It shares this habitat with several other
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
species, including the Nigerian white-throated guenon (''Cercopithecus erythrogaster pococki''), the
red-capped mangabey The collared mangabey (''Cercocebus torquatus''), also called red-capped mangabey and white-collared mangabey is a species of primate in the family Cercopithecidae of Old World monkeys. It formerly included the sooty mangabey as a subspecies. ...
(''Cercocebus torquatus''), the
putty-nosed monkey The greater spot-nosed monkey or putty-nosed monkey (''Cercopithecus nictitans'') is one of the smallest Old World monkeys. It is a guenon of the '' C. mitis'' group, native to West Africa and living to some extent in rain forests, but more often ...
(''Cercopithecus nictitans''), the
mona monkey The mona monkey (''Cercopithecus mona'') is an Old World monkey that lives in western Africa between Ghana and Cameroon. The mona monkey can also be found on the island of Grenada as it was transported to the island aboard slave ships headed to ...
(''Cercopithecus mona''), and maybe also the
olive colobus The olive colobus monkey (''Procolobus verus''), also known as the green colobus or Van Beneden's colobus, is a species of primate in the family Cercopithecidae. Its English name refers to its dull olive upperparts. It is the smallest example ...
(''Procolobus verus'').


Status

When it was first discovered, the Niger Delta red colobus was locally common but under some pressure from deforestation, especially logging of ''
Hallea ledermannii ''Mitragyna stipulosa'' is a species of flowering plant in the family Rubiaceae. It is found in Angola, Cameroon, Central African Republic, the Republic of the Congo, the Democratic Republic of the Congo, Gabon, Gambia, Ghana, Guinea, Nigeria, S ...
'', which is an important food tree for the monkey. Since then, pressure from
bushmeat Bushmeat is meat from wildlife species that are hunted for human consumption, most often referring to the meat of game in Africa. Bushmeat represents a primary source of animal protein and a cash-earning commodity for inhabitants of humid tropi ...
hunting and logging has increased. Red colobus monkeys generally appear to be particularly sensitive to hunting and habitat disturbance, hence concerns that the species may be on the verge of extinction. The Niger Delta red colobus was still found in the
Edumanom Forest Reserve The Edumanom Forest Reserve is an area in the Niger Delta region of South East Nigeria, that is home to some of the last chimpanzees in Nigeria. It covers part of the old Nembe Kingdom, which is now divided into the Nembe and Brass local government ...
in 2008. However, as of 2008, the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN) considers it critically endangered, due a reduction in population of more than 80% over the past 30 years largely caused by hunting and habitat loss. In 2010, the Niger Delta red colobus was included in the list of The World’s 25 Most Endangered Primates, published by the IUCN and other organizations.


References

{{Authority control
Niger Delta red colobus The Niger Delta red colobus (''Piliocolobus epieni'') is a critically endangered species of colobus monkey endemic to the western part of the Niger Delta. It is threatened by hunting and habitat loss. Taxonomy From the time it first became kno ...
Niger Delta red colobus The Niger Delta red colobus (''Piliocolobus epieni'') is a critically endangered species of colobus monkey endemic to the western part of the Niger Delta. It is threatened by hunting and habitat loss. Taxonomy From the time it first became kno ...
Niger Delta red colobus The Niger Delta red colobus (''Piliocolobus epieni'') is a critically endangered species of colobus monkey endemic to the western part of the Niger Delta. It is threatened by hunting and habitat loss. Taxonomy From the time it first became kno ...
Niger Delta red colobus The Niger Delta red colobus (''Piliocolobus epieni'') is a critically endangered species of colobus monkey endemic to the western part of the Niger Delta. It is threatened by hunting and habitat loss. Taxonomy From the time it first became kno ...
Niger Delta red colobus The Niger Delta red colobus (''Piliocolobus epieni'') is a critically endangered species of colobus monkey endemic to the western part of the Niger Delta. It is threatened by hunting and habitat loss. Taxonomy From the time it first became kno ...
Critically endangered fauna of Africa
Niger Delta red colobus The Niger Delta red colobus (''Piliocolobus epieni'') is a critically endangered species of colobus monkey endemic to the western part of the Niger Delta. It is threatened by hunting and habitat loss. Taxonomy From the time it first became kno ...
Primates of Africa Taxa named by Peter Grubb