Netta Caryophyllacea
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The pink-headed duck (''Rhodonessa caryophyllacea'') is a large diving duck that was once found in parts of the Gangetic plains of India, parts of Maharashtra, Bangladesh and in the riverine swamps of Myanmar but feared extinct since the 1950s. Numerous searches have failed to provide any proof of continued existence. It has been suggested that it may exist in the inaccessible swamp regions of northern Myanmar and some sight reports from that region have led to its status being declared as "Critically Endangered" rather than extinct. The genus placement has been disputed and while some have suggested that it is close to the red-crested pochard ('' Netta rufina''), others have placed it in a separate genus of its own. It is unique in the pink colouration of the head combined with a dark body. A prominent wing patch and the long slender neck are features shared with the common Indian spot-billed duck. The eggs have also been held as particularly peculiar in being nearly spherical.


Description

The male pink-headed duck is unmistakable when a good view is had. Both sexes are 41–43 cm and long-billed with long necks and peaked heads. The male has a pink bill, head and neck while the female has a pale pinkish head and neck with a paler bill. The black of the body extends as a narrow strip on the front of the neck. Wings have a leading white edge. In flight it would not contrast as much as the syntopic white-winged duck. Wing does not have the dark trailing edge of the red-crested pochard. Confusion with male red-crested pochards stems mainly from observations of swimming birds, as the latter species also has a conspicuous red head (although the color is actually very different from the pink-headed duck). Indian spot-billed ducks, on the other hand, can look similar to female pink-headed ducks when in flight and seen from a distance, and if seen from behind, they could be mistaken for males too. The upper side of the wing is distinguishing, with dark green secondaries (speculum) and prominent white tertiaries in the spot-billed duck and a pinkish-beige speculum, much lighter than its surroundings, in the pink-headed duck. If the upper part of the wings cannot be reliably seen, they are all but indistinguishable except to expert observers in good visibility conditions. Young birds had a nearly whitish head without a trace of pink and a mellow two note call ''wugh-ah'' has been attributed to the species. Its breeding habitat is lowland marshes and pools in tall-grass jungle. The nest is built amongst grass. The eggs, six or seven in a clutch, are very spherical and creamy white. The eggs measure 1.71 to 1.82 inches long and 1.61 to 1.7 inches wide. They were believed to have been non-migratory and found singly or in pairs and very rarely in small groups. Pink-headed ducks are believed to have eaten water plants and molluscs. Like '' Netta'' species, they typically up-ended or dabbled for food and did not dive like a pochard.


Distribution

Allan Octavian Hume and Stuart Baker noted that the stronghold of the species was north of the Ganges and west of the Brahmaputra, mainly in Maldah, Purnea, Madhubani and Purulia districts of present-day Bihar. It was said to be commoner in Singhbum. Hume collected a specimen in Manipur which he noted was very rare, hiding among dense reeds in Loktak Lake.
Edward Blyth Edward Blyth (23 December 1810 – 27 December 1873) was an English zoologist who worked for most of his life in India as a curator of zoology at the museum of the Asiatic Society of India in Calcutta. Blyth was born in London in 1810. In 1841 ...
claimed that it was found in the Rakhine state of Burma. Brian Houghton Hodgson obtained specimens from Nepal. A few records were also noted from Delhi, Sindh and Punjab. The populations (possibly) undertook local seasonal movements, resulting in scattered historic records as far as Punjab, Maharahtra and Andhra Pradesh. Thus, resulting in rare sightings of the species in Maharashtra. Birds were also reported from the Oudh region some from very close to Lucknow. Specimens were shot at Najafgarh lake in the Delhi district. Jerdon obtained specimens of the bird from further south although he did not personally observe any in the wild until he visited Bengal.


Taxonomy and systematics

The pink-headed duck was described by John Latham in 1790 under the genus ''Anas''. In describing the species, it is possible that he made of use of a painting in the collection of Lady Impey, wife of Sir
Elijah Impey Sir Elijah Impey (13 June 17321 October 1809) was a British judge, the first chief justice of the Supreme Court of Judicature at Fort William in Bengal, Chief Justice of the Sadr Diwani Adalat and MP for New Romney. Life He was born the young ...
who was Chief Justice of court in Calcutta from 1774 to 1783. Mary Impey maintained a menagerie in Calcutta and commissioned Indian artists such as Bhawani Das from Patna to illustrate animals in the collection. The Impeys moved to England, and after the death of her husband, she sold these paintings at auction in 1810. Some of them were acquired by the
13th Earl of Derby Edward Smith-Stanley, 13th Earl of Derby (21 April 1775 – 30 June 1851), KG, of Knowsley Hall in Lancashire (styled Lord Stanley from 1776 to 1832, known as Baron Stanley of Bickerstaffe from 1832-4), was a politician, peer, landowner, bui ...
. The genus ''Rhodonessa'' was originally created for this species alone.
Jean Delacour Jean may refer to: People * Jean (female given name) * Jean (male given name) * Jean (surname) Fictional characters * Jean Grey, a Marvel Comics character * Jean Valjean, fictional character in novel ''Les Misérables'' and its adaptations * Jea ...
and
Ernst Mayr Ernst Walter Mayr (; 5 July 1904 – 3 February 2005) was one of the 20th century's leading evolutionary biologists. He was also a renowned Taxonomy (biology), taxonomist, tropical explorer, ornithologist, Philosophy of biology, philosopher o ...
, in their 1945 revision of the family Anatidae considered it a somewhat abnormal member of the ''Anatini'' (or river-ducks) group because the hind toe is slightly lobed, display behaviour and the tendency to feed at the surface. The birds were observed in European aviaries and although they never bred, the males displayed often and this involved puffing the neck feathers, lowering the neck to rest on the back and then stretching up the neck while producing a wheezy whistle like a
mallard The mallard () or wild duck (''Anas platyrhynchos'') is a dabbling duck that breeds throughout the temperate and subtropical Americas, Eurasia, and North Africa, and has been introduced to New Zealand, Australia, Peru, Brazil, Uruguay, Arge ...
. A study of its tracheal anatomy by Alfred Henry Garrod in 1875 suggested that it had a "slight fusiform dilatation" in the anterior syringeal region. The "bulba ossea" at the lower part of the male syrinx is peculiar in being swollen. The colour pattern has also been considered unique, lacking any of the metallic colours on the secondaries that are characteristic of the Anatini. The other unique feature being the somewhat large and nearly spherical shape of the eggs. All of these features supporting the retention of the species in a separate genus. Such mid-tracheal swellings were found only in Mergini and Aythyini and is extremely rare in the genus '' Anas''. This tracheal bulla is rounded in ''Anas'' but angular with fenestrae in '' Netta'' and '' Aythya''. Johnsgard considered '' Marmaronetta'' and ''Rhodonessa'' as intermediate in form. Based on the available morphological and behavioural evidence, especially the structure of the
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
and the structure of tracheal rings,
Sidney Dillon Ripley Sidney Dillon Ripley II (September 20, 1913 – March 12, 2001) was an American ornithologist and wildlife conservationist. He served as secretary of the Smithsonian Institution for 20 years, from 1964 to 1984, leading the institution through ...
suggested that it was undoubtedly in the Aythyini. A study found that ''Rhodonessa'' was closely allied to the red-crested pochard (''Netta rufina'') suggesting that the two species be placed in the same genus. ''Rhodonessa'' was described prior to ''Netta'' which would then make ''Rhodonessa rufina'' the name of choice, however these changes have not been widely accepted. The pink colour is derived from a carotenoid pigment which is unusual among ducks and known only from a few other species such as the pink-eared duck which are not closely related.


Status

This duck formerly occurred in eastern India, Bangladesh and northern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, but is now probably
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
. It was always rare, and the last confirmed sighting, by C. M. Inglis, was from Bhagownie, Darbhangha District, in June 1935, with reports from India persisting until the early 1960s. These include reports from
Monghyr Munger, formerly spelt as Monghyr, is a twin city and a Municipal Corporation situated in the Indian state of Bihar. It is the administrative headquarters of Munger district and Munger Division. Munger was one of the major cities in Eastern In ...
and from near Shimla.
Sidney Dillon Ripley Sidney Dillon Ripley II (September 20, 1913 – March 12, 2001) was an American ornithologist and wildlife conservationist. He served as secretary of the Smithsonian Institution for 20 years, from 1964 to 1984, leading the institution through ...
considered it likely extinct in 1950. In 1988,
Rory Nugent Rory Nugent (born 1952) is an American explorer and writer. He was based out of New Bedford, Massachusetts from 1988 to 2004. Biography Nugent was born in New York. After he graduated from Williams College in 1975, he went to sea aboard freighters ...
, an American birder, and Shankar Barua of Delhi, reported spotting the elusive bird on the banks of the Brahmaputra. The pair started their quest for the bird at Saikhoa ghat on the north-eastern end of the river on the Indian side of the border. After 29 days of sailing, Nugent said that he saw the pink-headed duck amidst a flock of other waterbirds. However, Nugent and Barua's claimed sighting has not been widely accepted. Reports of pink-headed ducks after the 1960s have been received from the largely unexplored Mali Hka and Chindwin Myit drainages in Northern Myanmar. While the area is not very well surveyed by scientists, searches have been inconclusive and confusion with the red-crested pochard and the Indian spot-billed duck has been a common source of supposed pink-headed duck sightings. A report on a survey in the Hu Kaung valley in November 2003 concluded that there is sufficient reason to believe that pink-headed ducks may still exist in Northern Myanmar's
Kachin State Kachin State ( my, ကချင်ပြည်နယ်; Kachin: ), also known by the endonym Kachinland, is the northernmost state of Myanmar. It is bordered by China to the north and east (Tibet and Yunnan, specifically and respectively); Sh ...
, but a thorough survey of the Nat Kaung river between Kamaing and Shadusup in October 2005 failed to find this species; a number of interesting ducks were observed, but they turned out to be Indian spot-billed ducks or white-winged ducks. Suggestions have been made that it may be nocturnal. In 2017, an expedition to find the species by Global Wildlife Conservation also failed, with evidence indicating that the biodiversity in the general area around
Indawgyi Lake Indawgyi Lake ( ) is one of the largest inland lakes in Southeast Asia. It is located in Mohnyin Township in the Kachin State of Myanmar. The lake measures east to west, and north to south. There are over 20 villages around the lake. The predomi ...
and its surrounding areas was heavily declining due to habitat degradation. Anecdotes from residents in the area, however, indicate that the bird may have lived in the area far more recently than the last confirmed report from 1910, possibly as recently as 2010. One resident stated that a pink-headed duck was sighted in 1998, associating with a flock of
gadwall The gadwall (''Mareca strepera'') is a common and widespread dabbling duck in the family Anatidae. Taxonomy The gadwall was first described by Carl Linnaeus in his landmark 1758 10th edition of ''Systema Naturae''. DNA studies have shown that ...
and pintail. Another, more dubious report stated that shortly after a failed expedition in the area by
Birdlife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
ended, a local hunter caught a live male and a female or juvenile pink-headed duck, and contacted Myanmar's Biodiversity and Nature Conservation Association to sell it to them for a fee. The offer was declined, and the hunter killed both ducks. Another hunter recalled that when the habitat was in good condition, pink-headed ducks were regulars in the area, possibly up to 2014. They were apparently most common during February, and he also could mimic their possible calls, though it is unknown whether these calls were truly by pink-headed ducks. The hunter also said that there were large, impassable ponds in the wetland's center that may still hold pink-headed ducks, but these could only be accessed with a drone, which are banned in the region. The reason for its disappearance was probably habitat destruction. It is not known why it was always considered rare, but the rarity is believed to be genuine (and not an artefact of insufficient fieldwork) as its erstwhile habitat was frequently scoured by hunters in Colonial times. The pink-headed duck was much sought after by hunters and later as an ornamental bird, mainly because of its unusual plumage. Like most diving ducks, it was not considered good eating, which should facilitate the survival of any remnant birds. The last specimen was obtained in 1935 in
Darbhanga Darbhanga is the fifth-largest city and municipal corporation in the Indian state of Bihar situated centrally in Mithila region. Darbhanga is the headquarters of the Darbhanga district and the Darbhanga division. It was the seat of the erstw ...
, Andhra Pradesh, India by C. M. Inglis. Some birds were also kept in the aviaries of
Jean Théodore Delacour Jean Théodore Delacour (26 September 1890 – 5 November 1985) was a French ornithologist and aviculturist. He later became American. He was renowned for not only discovering but also rearing some of the rarest birds in the world. He establish ...
in
Clères Clères () is a commune in the Seine-Maritime department in the Normandy region in north-western France. Geography A farming and forestry market town situated by the banks of the Clérette river, in the Pays de Caux, some northeast of Rouen, ...
( France) and
Alfred Ezra Alfred "Chips" Ezra, OBE (1872 – 1 August 1955) was a British breeder and keeper of birds. He built up a collection of rare birds at Foxwarren Park in southern England that was considered the finest of its kind. Biography Alfred Ezra was bor ...
at Foxwarren Park ( England) where the last known birds lived in captivity. The only known photographs of the species were taken here and include one of a pair taken around 1925 by
David Seth-Smith David Seth-Smith FZS, MBOU (9 March 1875 – 30 October 1963) was a British zoologist, wildlife artist, nature broadcaster and author. His career included spells as Curator of Mammals and Birds for the Zoological Society of London and ed ...
.


References


Other sources

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External links


BirdLife International Species Factsheet
{{Authority control pink-headed duck pink-headed duck Birds of India Birds of Myanmar Birds of Bangladesh pink-headed duck pink-headed duck Articles containing video clips