Nerodia Fasciata
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The banded water snake or southern water snake (''Nerodia fasciata'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of mostly aquatic, nonvenomous,
colubrid Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ever ...
snakes
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the
Midwest The Midwestern United States, also referred to as the Midwest or the American Midwest, is one of four Census Bureau Region, census regions of the United States Census Bureau (also known as "Region 2"). It occupies the northern central part of ...
and Southeastern United States.


Geographic range

''N. fasciata'' is natively found from
Indiana Indiana () is a U.S. state in the Midwestern United States. It is the 38th-largest by area and the 17th-most populous of the 50 States. Its capital and largest city is Indianapolis. Indiana was admitted to the United States as the 19th s ...
, south to
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
, and east to Florida. In 1992, its congener '' Nerodia sipedon'' (northern or common water snake) and it were found in three sites in California by the
US Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
(USFWS). In 2009, more than 300 banded water snakes were caught in suburbs of Los Angeles by the Nerodia Working Group of USFWS. Then in May 2016, the species was found in the
Colorado River The Colorado River ( es, Río Colorado) is one of the principal rivers (along with the Rio Grande) in the Southwestern United States and northern Mexico. The river drains an expansive, arid drainage basin, watershed that encompasses parts of ...
basin near
Yuma, Arizona Yuma ( coc, Yuum) is a city in and the county seat of Yuma County, Arizona, United States. The city's population was 93,064 at the 2010 census, up from the 2000 census population of 77,515. Yuma is the principal city of the Yuma, Arizona, M ...
. Further trapping did indeed catch large numbers of them, indicating that a thriving invasive population exists in that area.


Description

Adults of the banded water snake measure from in total length, with a record size (in the Florida subspecies) of in total length. In one study, the average body mass of adult banded water snakes was . It is typically gray, greenish-gray, or brown in color, with dark crossbanding. Many specimens are so dark in color that their patterning is barely discernible. The ventrum (belly) is typically an off-white to white. They have flat heads, and are fairly heavy-bodied. Banded water snakes may also be identified by a dark stripe which extends from the eye to the angle of the jaw. If irritated, they release a foul-smelling musk to deter predators. This species also exhibits sexual dimorphism in which the female is generally longer and heavier than the male. Their appearance leads them to be frequently mistaken for other snakes with which they share a habitat, including the venomous
cottonmouth ''Agkistrodon piscivorus'' is a species of pit viper in the subfamily Crotalinae of the family Viperidae. It is one of the world's few semiaquatic vipers (along with the Florida cottonmouth), and is native to the southeastern United States. As ...
.


Habitat

''N. fasciata'' inhabits most freshwater environments such as lakes,
marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
es, ponds, and streams. Conant, R. 1975. ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition.'' Houghton Mifflin. Boston. xviii + 429 pp. (paperback). (''Natrix fasciata fasciata'', p. 146 + Plate 20 + Map 100.) Banded water snakes are active both day and night and may be seen basking on logs or branches overhanging the water or foraging in shallow water. They will typically stay within emergent vegetation along the shoreline or in the shallow littoral zone of their habitat.


Diet

The species preys mainly on fish and frogs. Juveniles mainly consume fish and shift towards eating frogs as adults. This shift may be caused by large frogs being mechanically too difficult for juveniles to consume (which may suggest juveniles may be unable to open their jaws wide enough for adult frogs), because large frogs are energetically too costly for juveniles to catch, or because juveniles are at risk if swallowing prey requires a long time. Using its
vomeronasal organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. ...
, also called Jacobson's organ, the snake can detect
parvalbumin Parvalbumin (PV) is a calcium-binding protein with low molecular weight (typically 9-11 kDa). In humans, it is encoded by the ''PVALB'' gene. It is not a member of the albumin family; it is named for its size (''parv-'', from Latin ''parvus'' smal ...
s in the
cutaneous Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other animal coverings, such as the arthropod exoskeleton, have different de ...
mucus Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It is ...
of its prey.


Reproduction

The species is viviparous, giving birth to live young. The brood size varies from 15 to 20 young born in late July or August. Newborns are 200–240 mm (about 8.0–9.5 in) in total length. Wright, A.H., and A.A. Wright. 1957. ''Handbook of Snakes of the United States and Canada.'' Comstock. Ithaca and London. 1,105 pp. (in 2 volumes) (''Natrix sipedon fasciata'', pp. 525–529, Figure 156.) The banded water snake is able to hybridize with the
common watersnake The common watersnake (''Nerodia sipedon'') is a species of large, nonvenomous, common snake in the family Colubridae. The species is native to North America. It is frequently mistaken for the venomous cottonmouth (''Agkistrodon piscivorus''). C ...
, although this is not frequent. Physical characteristics are insufficient to distinguish such hybrids, and DNA analysis is required.


Subspecies

The three recognized
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of ''N. fasciata'', including the
nominotypical subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, are: *''N. f. confluens'' (
Blanchard Blanchard is a French family name. It is also used as a given name. It derives from the Old French word ''blanchart'' which meant "whitish, bordering upon white". It is also an obsolete term for a white horse. Geographical distribution As of 2014, ...
, 1923)
- Broad-banded water snake -
Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...
,
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...
,
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
,
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
,
Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ...
,
East Texas East Texas is a broadly defined cultural, geographic, and ecological region in the eastern part of the U.S. state of Texas that comprises most of 41 counties. It is primarily divided into Northeast and Southeast Texas. Most of the region consi ...
. *''N. f. fasciata'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1766)
-
East Texas East Texas is a broadly defined cultural, geographic, and ecological region in the eastern part of the U.S. state of Texas that comprises most of 41 counties. It is primarily divided into Northeast and Southeast Texas. Most of the region consi ...
,
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
,
southeastern Oklahoma Choctaw Country is the Oklahoma Department of Tourism and Recreation's official tourism designation for Southeastern Oklahoma. The name was previously Kiamichi Country until changed in honor of the Choctaw Nation headquartered there. The current ...
,
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...
, western
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
,
southern Alabama South Alabama is the various parts of southern Alabama. Although it is not a strictly defined geographic region, it generally includes all Alabama counties south of the Black Belt. In that view, ''South Alabama'' consists of just the two counties ...
,
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, southern
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
,
South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...
,
North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...
, southeast
Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ...
and
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolita ...
. *'' N. f. pictiventris'' (
Cope The cope (known in Latin as ''pluviale'' 'rain coat' or ''cappa'' 'cape') is a liturgical vestment, more precisely a long mantle or cloak, open in front and fastened at the breast with a band or clasp. It may be of any liturgical colours, litu ...
, 1895)
– Florida banded water snake -
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
,
southeast Georgia Southeast Georgia is a nineteen-county region located south of the Altamaha River in the U.S. state of Georgia. The 2010 census calculated the region's population to be at 523,905. The largest city in this region is Valdosta and the largest count ...
. Introduced to
Brownsville, Texas Brownsville () is a city in Cameron County in the U.S. state of Texas. It is on the western Gulf Coast in South Texas, adjacent to the border with Matamoros, Mexico. The city covers , and has a population of 186,738 as of the 2020 census. It ...
and
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
.


Taxonomy

Some sources consider '' N. clarkii compressicauda'' and ''N. clarkii taeniata'' to be
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of ''N. fasciata''. Also, some sources have considered ''Nerodia fasciata'' to be a subspecies of '' Nerodia sipedon''. Schmidt, K.P., and D.D. Davis. 1941. ''Field Book of Snakes of the United States and Canada.'' G.P. Putnam's Sons. New York. 365 pp. (''Natrix sipedon fasciata'', pp. 221–222, Figure 72. + Plate 24, Center, on p. 344.)


References


Other sources


Herps of Texas: ''Nerodia fasciata''


External links

*
California Nerodia Watch – iNaturalist


Further reading

* Linnaeus, C. 1766. ''Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, diferentiis, synonymis, locis. Tomus I. Editio Duodecima, Reformata.'' L. Salvius. Stockholm. 532 pp. (''Coluber fasciatus'', p. 378.) {{Authority control Nerodia Articles containing video clips Extant Pleistocene first appearances Reptiles described in 1766 Taxa named by Carl Linnaeus Reptiles of the United States Fauna of the Southeastern United States