Nepenthes Flava
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''Nepenthes flava'' is a tropical
pitcher plant Pitcher plants are several different carnivorous plants which have modified leaves known as pitfall traps—a prey-trapping mechanism featuring a deep cavity filled with digestive liquid. The traps of what are considered to be "true" pitcher p ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to northern
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
, where it grows in
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
at 1800–2200 m
above sea level Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The comb ...
.McPherson, S.R. 2009. ''
Pitcher Plants of the Old World ''Pitcher Plants of the Old World'' is a two-volume monograph by Stewart McPherson on the pitcher plants of the genera ''Nepenthes'' and ''Cephalotus''. It was published in May 2009 by Redfern Natural History Productions and covers all species kno ...
''. 2 volumes. Redfern Natural History Productions, Poole.
The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''flava'' is derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
word for "yellow" and refers to the typical colouration of the plant's upper pitchers and other vegetative parts.


Botanical history

''Nepenthes flava'' entered cultivation several years prior to being described and was known from at least 2004 under the placeholder name "''Nepenthes'' spec. nov. Sumatra".Wistuba, A. 2004
''Nepenthes'' spec. nov. Sumatra
Wistuba.com. rchived page from April 15, 2004/ref> The species was formally described by
Andreas Wistuba Andreas Wistuba (born 4 March 1967) is a German taxonomist and botanist specialising in the carnivorous plant genera ''Heliamphora'' and '' Nepenthes''. More than half of all known ''Heliamphora'' species have been described by Wistuba. Pub ...
,
Joachim Nerz Dr. Joachim Nerz (born 1964) is a German taxonomist and botanist specialising in the carnivorous plant genera ''Heliamphora'' and '' Nepenthes''. Nerz has described several new species, mostly with Andreas Wistuba. Publications * Schlauer, J. ...
and
Andreas Fleischmann Andreas ( el, Ἀνδρέας) is a name usually given to males in Austria, Greece, Cyprus, Denmark, Armenia, Estonia, Ethiopia, Eritrea, Finland, Flanders, Germany, Norway, Sweden, Switzerland, Romania, the Netherlands, and Indonesia. The name ...
in an issue of ''
Blumea ''Blumea'' is a genus of flowering plants of the family Asteraceae. Characteristics Genus ''Blumea'' is found in the tropical and sub-tropical zones of Asia, especially the Indian Subcontinent and Southeast Asia. A few species are found in Aust ...
'' published on July 4, 2007.''Nepenthes flava'' Wistuba, Nerz & A.Fleischm.
International Plant Names Index The International Plant Names Index (IPNI) describes itself as "a database of the names and associated basic bibliographical details of seed plants, ferns and lycophytes." Coverage of plant names is best at the rank of species and genus. It inclu ...
(IPNI).
The description was based on cultivated plant material; ''Wistuba cult. Wistuba 100201'' was designated as the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
. This specimen was collected at an altitude of 1800 m and is deposited at the
National Herbarium of the Netherlands National may refer to: Common uses * Nation or country ** Nationality – a ''national'' is a person who is subject to a nation, regardless of whether the person has full rights as a citizen Places in the United States * National, Maryland, ce ...
(L) in
Leiden Leiden (; in English and archaic Dutch also Leyden) is a city and municipality in the province of South Holland, Netherlands. The municipality of Leiden has a population of 119,713, but the city forms one densely connected agglomeration wit ...
, together with an isotype. The describing authors explained their choice of the
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''flava'' as follows:
The specific epithet ‘flava’ refers to the bright yellow colour of the upper pitchers and most of the lower pitchers. Climbing plants especially give the impression of a mainly yellow plant.
A detailed and slightly modified description of ''N. flava'' appeared in Stewart McPherson's 2009 monograph, ''
Pitcher Plants of the Old World ''Pitcher Plants of the Old World'' is a two-volume monograph by Stewart McPherson on the pitcher plants of the genera ''Nepenthes'' and ''Cephalotus''. It was published in May 2009 by Redfern Natural History Productions and covers all species kno ...
''. Whereas the type description mentions a basal crest on the underside of the lid, McPherson writes that no appendages are present in either lower or upper pitchers.


Description

''Nepenthes flava'' is a climbing plant growing to a height of 6 m. It only remains in the rosette stage for a short time before transitioning into a scrambling
vine A vine (Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners themselv ...
. The
stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...
, which may be branched, is around 3 mm in diameter and has roughly cylindrical internodes measuring up to 14 cm in length. The stem ranges in colour from green to dark red. The
lamina Lamina may refer to: Science and technology * Planar lamina, a two-dimensional planar closed surface with mass and density, in mathematics * Laminar flow, (or streamline flow) occurs when a fluid flows in parallel layers, with no disruption betwee ...
(leaf blade) is linear, oblong, or narrowly
obovate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
, and measures up to 9 cm in length by 2.5 cm in width. Its apex is usually acute, but may also be obtuse. The base of the lamina is
attenuate In physics, attenuation (in some contexts, extinction) is the gradual loss of flux intensity through a medium. For instance, dark glasses attenuate sunlight, lead attenuates X-rays, and water and air attenuate both light and sound at variable a ...
and clasps the stem for around half to three-quarters of its circumference. Three prominent longitudinal veins are present on either side of the
midrib This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
. Indistinct
reticulate Reticulation is a net-like pattern, arrangement, or structure. Reticulation or Reticulated may refer to: * Reticulation (single-access key), a structure of an identification tree, where there are several possible routes to a correct identificatio ...
veins are also present. The lamina is yellowish-green in colour and the midrib may be reddish.
Tendril In botany, a tendril is a specialized stem, leaf or petiole with a threadlike shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as ''Cuscuta''. There are many plants that have tendr ...
s reach 24 cm in length and may be green to red. Rosette and lower pitchers are either
infundibular An infundibulum (Latin for ''funnel''; plural, ''infundibula'') is a funnel-shaped cavity or organ. Anatomy * Brain: the pituitary stalk, also known as the ''infundibulum'' and ''infundibular stalk'', is the connection between the hypothalamus and ...
throughout or
ovate Ovate may refer to: *Ovate (egg-shaped) leaves, tepals, or other botanical parts *Ovate, a type of prehistoric stone hand axe *Ovates, one of three ranks of membership in the Welsh Gorsedd *Vates In modern English, the nouns vates () and ovat ...
and variably inflated. They are small, measuring only up to 7 cm in height by 4 cm in width. A pair of wings (≤3 mm wide) often runs down the upper third of the pitcher's ventral surface, bearing fringe elements around 5 mm long. Occasionally, the wings may be absent altogether. The
peristome Peristome (from the Greek ''peri'', meaning 'around' or 'about', and ''stoma'', 'mouth') is an anatomical feature that surrounds an opening to an organ or structure. Some plants, fungi, and shelled gastropods have peristomes. In mosses In mosses, ...
varies from cylindrical to slightly expanded and has a wavy outer margin. It is up to 12 mm wide and bears fine ribs up to 0.6 mm high and spaced up to 1 mm apart. On the inner margin of the peristome, these ribs terminate in teeth up to 1.5 mm long, with the largest located towards the rear. The pitcher mouth is round and slightly elongated towards the rear, although it rarely exhibits a neck. The pitcher lid or operculum is narrowly ovate or elliptic and measures up to 3.5 cm in length by 2 cm in width. On its lower surface, the lid bears a basal
gland In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...
ular crest or no appendages at all, as well as numerous circular
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
glands (0.3 mm in diameter), which are concentrated around the midline and crest. An unbranched,
filiform Filiform, thread or filament like, can refer to: * Filiform, a common term used in botany to describe a thread-like shape *Filiform, or filiform catheter In medicine, a catheter (/ˈkæθətər/) is a thin tubing (material), tube made from m ...
spur A spur is a metal tool designed to be worn in pairs on the heels of riding boots for the purpose of directing a horse or other animal to move forward or laterally while riding. It is usually used to refine the riding aids (commands) and to back ...
(≤4 mm long) is inserted near the base of the lid. Unlike their aerial counterparts, lower pitchers vary greatly in pigmentation. The pitcher cup may be yellow, orange, red, or even purple, whereas the inner surface may be light yellow, light orange, or whitish. The peristome is usually darker than the rest of the pitcher, typically being dark orange to purple, although it may be creamy white and occasionally exhibits red stripes. The lid has a similar colour to the pitcher cup, usually having a darker upper surface with dark red speckles. Upper pitchers are tubular to narrowly infundibular in the lower two-thirds, becoming broadly infundibular above. Characteristically, the hollow pitcher tube often continues past the curved basal portion and for some distance up the tendril. Upper pitchers are similar in size to their terrestrial counterparts, reaching 6 cm in height by 3.5 cm in width. The peristome is flattened and expanded, reaching 15 mm in width. As in lower pitchers, its outer margin is undulate. The peristome ribs are up to 0.2 mm high and spaced up to 0.5 mm apart. The lid is ovate or elliptic and has a somewhat
truncate In mathematics and computer science, truncation is limiting the number of digits right of the decimal point. Truncation and floor function Truncation of positive real numbers can be done using the floor function. Given a number x \in \mathbb ...
apex. It measures up to 4 cm in length by 2.5 cm in width. A reduced basal crest may be present on the underside of the lid or it may be absent entirely. Other parts are similar to those found in terrestrial traps. Upper pitchers are typically yellow throughout, but some specimens may have a completely red or red-striped peristome and mature traps may be orange or reddish on the upper surface of the lid. ''Nepenthes flava'' has a
racemose A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
up to 15 cm long. Male inflorescences are more
floriferous {{Short pages monitor