''Nepenthes bokorensis'' is a tropical

pitcher plant Pitcher plants are several different carnivorous plants which have modified leaves known as pitfall traps—a prey-trapping mechanism featuring a deep cavity filled with digestive liquid. The traps of what are considered to be "true" pitcher p ...

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...

. It is known from Mount Bokor (also ''Phnom Bokor'' or Bokor Hill) in the south of the country, and an as yet undetermined specimen suggests that it may also be present in other parts of the

Dâmrei Mountains The Dâmrei Mountains (literally the "Elephant Mountains", km, ភ្នំដំរី, Chuŏr Phnum Dâmrei), refer to a mountain range situated in south-western Cambodia, traversing around north-south as a succession of the Cardamom Mounta ...

Kampot Province Kampot ( km, កំពត ) is a province in southwestern Cambodia. It borders the provinces of Koh Kong and Kampong Speu to the north, Takéo to the east, Kep and the country of Vietnam ( Kiên Giang) to the south, and Sihanoukville to the ...

. The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''bokorensis'' refers to both Mount Bokor and

Bokor National Park Preah Monivong Bokor National Park ( km, ឧទ្យានជាតិព្រះមុនីវង្ស បូកគោ, ) is a national park in southern Cambodia's Kampot Province that was established in 1993 and covers . It is designated as ...

Botanical history

Despite only being formally described in 2009, ''N. bokorensis'' has been known since at least the early 20th century. The oldest known

herbarium A herbarium (plural: herbaria) is a collection of preserved plant specimens and associated data used for scientific study. The specimens may be whole plants or plant parts; these will usually be in dried form mounted on a sheet of paper (called ...

specimens are three taken by French

plant collector Plant collecting is the acquisition of plant specimens for the purposes of research, cultivation, or as a hobby. Plant specimens may be kept alive, but are more commonly dried and pressed to preserve the quality of the specimen. Plant collecting i ...

C. Geoffray on January 14, 1904. Two of these consist of lower pitchers with leaves, while the third comprises upper pitchers with leaves. All three specimens were collected from Popokvil falls, Mount Bokor, and are deposited at the

Museum National d'Histoire Naturelle The French National Museum of Natural History, known in French as the ' (abbreviation MNHN), is the national natural history museum of France and a ' of higher education part of Sorbonne Universities. The main museum, with four galleries, is loc ...

Paris Paris () is the capital and most populous city of France, with an estimated population of 2,165,423 residents in 2019 in an area of more than 105 km² (41 sq mi), making it the 30th most densely populated city in the world in 2020. S ...

. The next known collection of this species was made by J. E. Vidal in 1965. It consists of a plant with upper pitchers. Three years later, Marie Martin collected ''M. Martin 1231bis'', which includes lower pitchers and female floral material. Further herbarium material, consisting of two rosette plants, was collected by David Middleton and Meng Monyrak in 2001. In July 2007, François Sockhom Mey found ''N. bokorensis'' on Mount Bokor during a field trip to southern

. He posted photographs of the species on a number of online

message board An Internet forum, or message board, is an online discussion site where people can hold conversations in the form of posted messages. They differ from chat rooms in that messages are often longer than one line of text, and are at least temporar ...

s the following month. Mey formally described the species in the March 2009 issue of ''

Carniflora Australis ''Carniflora Australis'' was a biannual English-language periodical and the official publication of the Australasian Carnivorous Plant Society.Rice, B. 2010Carnivorous Plant Society Archives The Carnivorous Plant FAQ. rchived page from October 1 ...

'', the journal of the Australasian Carnivorous Plant Society. ''M. Martin 1231bis'', deposited at the Museum National d'Histoire Naturelle, was designated as the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

. A specimen collected by

Auguste Jean Baptiste Chevalier Auguste may refer to: People Surname * Arsène Auguste (born 1951), Haitian footballer * Donna Auguste (born 1958), African-American businesswoman * Georges Auguste (born 1933), Haitian painter * Henri Auguste (1759–1816), Parisian gold and ...

in 1917 may also represent this species and if confirmed as such would increase the known range of ''N. bokorensis'' to include other parts of the

. Material deposited at Forest Herbarium,

Bangkok Bangkok, officially known in Thai language, Thai as Krung Thep Maha Nakhon and colloquially as Krung Thep, is the capital and most populous city of Thailand. The city occupies in the Chao Phraya River delta in central Thailand and has an estima ...

(BKF) and identified as '' N. thorelii'' likely also belongs to ''N. bokorensis''. ''Nepenthes bokor'', described by

Martin Cheek Martin Roy Cheek (born 1960) is a botanist and taxonomist at the Royal Botanic Gardens, Kew.Alastair Robinson Alastair S. Robinson (born 1980) is a taxonomist and field botanist specialising in the carnivorous plant genus ''Nepenthes'', for which he is regarded as a world authority.Ellison, A. & Adamec, L. eds., 2017. Contributing Author Information ...

discovered two large new populations of ''N. bokorensis'' on the Mount Bokor

massif In geology, a massif ( or ) is a section of a planet's crust that is demarcated by faults or flexures. In the movement of the crust, a massif tends to retain its internal structure while being displaced as a whole. The term also refers to a ...

, which they called "Location B" and "Location C" (the original roadside population being "Location A").Mey, F.S. 2011
''Nepenthes bokorensis'' 2011 conservation update
''Strange Fruits: A Garden's Chronicle'', August 28, 2011.

Description

''Nepenthes bokorensis'' is a climbing plant, attaining a height of up to 7 m. In rosettes and on lower parts of the plant, the stem is up to 0.9 cm thick and circular in cross section. On climbing parts, the stem is

terete Terete is a term in botany used to describe a cross section that is circular, or like a distorted circle, with a single surface wrapping around it.Lichen Vocabulary, Lichens of North America Information, Sylvia and Stephen Sharnoff/ref> This is us ...

and measures up to 1 cm in diameter. Internodes are around 3 cm long. Nepenthes bokorensis plant

Leaves are

sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...

to sub-

petiolate Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, a ...

and

coriaceous This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...

(leathery) in texture. The

lamina Lamina may refer to: Science and technology * Planar lamina, a two-dimensional planar closed surface with mass and density, in mathematics * Laminar flow, (or streamline flow) occurs when a fluid flows in parallel layers, with no disruption betwee ...

or leaf blade is oblong to linear-

lanceolate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...

in shape and measures up to 35 cm in length by 8 cm in width. Its apex varies greatly, ranging from acute to obtuse and it may sometimes also be

acuminate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...

. The lamina is

attenuate In physics, attenuation (in some contexts, extinction) is the gradual loss of flux intensity through a medium. For instance, dark glasses attenuate sunlight, lead attenuates X-rays, and water and air attenuate both light and sound at variable a ...

at the base, clasping the stem by around three-quarters of its circumference and rarely becoming

decurrent ''Decurrent'' (sometimes decurring) is a term used in botany and mycology to describe plant or fungal parts that extend downward. In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petio ...

. Three longitudinal veins are present on either side of the

midrib This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...

. Pinnate veins originate obliquely from the midrib. Tendrils are up to 18 cm long and 2 mm in diameter. Those produced on upper parts of the stem may be coiled.

Rosette and lower pitchers measure up to 20 cm in height by 6 cm in width. They are

ovate Ovate may refer to: *Ovate (egg-shaped) leaves, tepals, or other botanical parts *Ovate, a type of prehistoric stone hand axe *Ovates, one of three ranks of membership in the Welsh Gorsedd *Vates In modern English, the nouns vates () and ovat ...

in the basal third, before narrowing to become cylindrical or slightly

infundibular An infundibulum (Latin for ''funnel''; plural, ''infundibula'') is a funnel-shaped cavity or organ. Anatomy * Brain: the pituitary stalk, also known as the ''infundibulum'' and ''infundibular stalk'', is the connection between the hypothalamus and ...

towards the peristome. A pair of fringed wings (≤15 mm wide) runs down the ventral surface of the pitcher cup. Fringe elements are up to 12 mm long. The pitcher mouth has an oblique insertion. The peristome is approximately cylindrical and may be up to 20 mm wide. It bears ribs (≤0.8 mm high) spaced up to 1 mm apart. The pitcher lid or operculum is

orbicular Orbicular is an adjective meaning "circular" Orbicular may also refer to: * Orbicular leaf, a plant leaf shape * Orbicularis oculi muscle, a muscle around the eye * Orbicularis oris muscle, a muscle around the mouth * Orbicular batfish, a specie ...

to broadly elliptic in shape, has a

cordate Cordate is an adjective meaning 'heart-shaped' and is most typically used for: * Cordate (leaf shape), in plants * Cordate axe, a prehistoric stone tool See also * Chordate A chordate () is an animal of the phylum Chordata (). All chordat ...

base, and measures up to 7 cm in length by 6 cm in width. The underside of the lid lacks appendages, but bears numerous, densely packed

crater Crater may refer to: Landforms *Impact crater, a depression caused by two celestial bodies impacting each other, such as a meteorite hitting a planet *Explosion crater, a hole formed in the ground produced by an explosion near or below the surfac ...

-like

gland In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...

s. Those concentrated along the midrib reach the greatest dimensions (≤0.5 mm in diameter), while the rest are much smaller (0.2 to 0.3 mm in diameter) and more sparsely distributed. A spur measuring up to 12 mm in length is inserted near the base of the lid. It is typically unbranched, but may also be

bifurcate Bifurcation or bifurcated may refer to: Science and technology * Bifurcation theory, the study of sudden changes in dynamical systems ** Bifurcation, of an incompressible flow, modeled by squeeze mapping the fluid flow * River bifurcation, the f ...

. Upper pitchers exhibit great variability in both form and colouration, but generally grow to 25 cm in height by 6 cm in width. They are infundibular throughout, becoming narrower in the basal portion. Wings are reduced to ribs in these aerial pitchers. The peristome is up to 17 mm wide and often somewhat flared at the margins. The peristome ribs are less developed than in lower pitchers, measuring only up to 0.4 mm in height, and spaced only up to 0.5 mm apart. The lid is similar to that found in terrestrial traps, although slightly smaller (up to 6 cm long by 5 cm wide) and often bearing a

revolute Revolute may mean: *in botany, having the edges rolled down or back; see Glossary of botanical terms#R **Revolute leaf *in engineering, being able to rotate but not slide ( of a joint) * "Revolute", a song by 12 Rods from ''Gay?'' See also * R ...

margin. Other parts of upper pitchers are similar to their lower counterparts. ''Nepenthes bokorensis'' has a racemose inflorescence measuring up to 100 cm in length. It bears up to 80 flowers borne on one-flowered

pedicels In botany, a pedicel is a stem that attaches a single flower to the inflorescence. Such inflorescences are described as ''pedicellate''. Description Pedicel refers to a structure connecting a single flower to its inflorescence. In the absenc ...

(≤9 mm long), or rarely two-flowered partial peduncles. In male plants, the peduncle reaches 70 cm and the rachis 30 cm, while female plants produce a rachis up to 20 cm long. Tepals are orbicular to elliptic, ranging in length from 2 mm in male flowers to 4 mm in female flowers. The former have

androphore The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s up to 2 mm long, while the latter bear ovaries around 4 mm long. Fruit are typically 10 to 25 mm long and each contain 50 to 100

fusiform Fusiform means having a spindle-like shape that is wide in the middle and tapers at both ends. It is similar to the lemon-shape, but often implies a focal broadening of a structure that continues from one or both ends, such as an aneurysm on a b ...

seeds measuring around 7 mm in length. All vegetative and floral parts of ''N. bokorensis'' bear an indumentum of silvery or brownish hairs up to 1 mm long. In some places this covering may be reduced, giving an almost glabrous appearance. It is not certain whether ''N. bokorensis'' produces a well-developed rootstock like other Indochinese species, but observations of cultivated plants suggest that this is likely to be the case.

Ecology

''Nepenthes bokorensis'' is known with certainty only from the Mount Bokor massif in

, where it has an altitudinal distribution of 800–1080 m

above sea level Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The comb ...

. Three populations are known from the mountain. An as yet undetermined specimen (''Aug. Chevalier 36411'') suggests that the species may also be present in other parts of the

The typical habitat of this species is seasonally dry upland

scrub Scrub(s) may refer to: * Scrub, low shrub and grass characteristic of scrubland * Scrubs (clothing), worn by medical staff * ''Scrubs'' (TV series), an American television program * Scrubs (occupation), also called "scrub tech," "scrub nurse," ...

. It also grows in stunted lower montane forest under sparse tree cover. Plants growing in exposed sites under direct sunlight are very stunted and often flower when less than 60 cm tall. The species attains much greater heights under the shade of surrounding vegetation. Natural hybrids with '' N. kampotiana'' have been recorded. In open areas, ''N. bokorensis'' is sometimes

sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species sh ...

with other

carnivorous plant Carnivorous plants are plants that derive some or most of their nutrients from trapping and consuming animals or protozoans Protozoa (singular: protozoan or protozoon; alternative plural: protozoans) are a group of single-celled eukaryot ...

s, including sundews and bladderworts.Mey, F.S. 2011
A stunning population of ''Drosera peltata'' on Phnom Bokor
''Strange Fruits: A Garden's Chronicle'', August 30, 2011. The ''N. bokorensis'' plants found by François Mey in 2007 grew alongside a crimson form of ''

Drosera peltata ''Drosera peltata'', commonly called the shield sundewSalmon, Bruce. 2001. ''Carnivorous Plants of New Zealand''. Ecosphere Publications. or pale sundew,Erickson, Rica. 1968. ''Plants of Prey in Australia''. Lamb Paterson Pty. Ltd.: Osborne Park, ...

'', but this population was destroyed by the time of the author's return in 2009. ''Drosera peltata'' still grows sympatrically with ''N. bokorensis'' at "Location C", a massive clearing discovered in 2011 on the Bokor plateau. The ''D. peltata'' plants at this site range in colour from yellowish-green to red and frequently exceed 30 cm in height. ''Nepenthes bokorensis'' also shares this habitat with another sundew—'' D. burmannii''—as well as with various grasses, stunded shrubs, and

lithophytic Lithophytes are plants that grow in or on rocks. They can be classified as either epilithic (or epipetric) or endolithic; epilithic lithophytes grow on the surfaces of rocks, while endolithic lithophytes grow in the crevices of rocks (and are als ...

orchids. Local bladderwort species include '' U. bifida'', '' U. odorata'', '' U. minutissima'', '' U. striatula'', '' U. subulata'', and '' U. uliginosa''. On Mount Bokor, the species grows on sandy,

acid In computer science, ACID ( atomicity, consistency, isolation, durability) is a set of properties of database transactions intended to guarantee data validity despite errors, power failures, and other mishaps. In the context of databases, a sequ ...

ic soils. The

substrate Substrate may refer to: Physical layers *Substrate (biology), the natural environment in which an organism lives, or the surface or medium on which an organism grows or is attached ** Substrate (locomotion), the surface over which an organism lo ...

has a pH of 4.6 according to the herbarium label of ''Middleton & Monyrak 589''.

Carnivory

The prey assemblage of ''N. bokorensis'' appears to consist primarily of ants.Hosoishi, S., S.-H. Park, S. Yamane & K. Ogata 2012
Species composition of ant prey of the pitcher plant ''Nepenthes bokorensis'' Mey (Nepenthaceae) in Phnom Bokor National Park, Cambodia.
''Cambodian Journal of Natural History'' 2012(1): 3–7. A 2012 study recorded 10 ant species, representing 9 genera and 3 subfamilies, from 30 ''N. bokorensis'' pitchers in

. ''Polyrhachis'' (''Myrma'') sp. was found to be the most abundant taxon, accounting for 40% of all specimens, followed by ''

Dolichoderus thoracicus ''Dolichoderus thoracicus'' is a species of ant in the genus '' Dolichoderus''. Described by Smith in 1860, the species is widespread in Asia.Shattuck, S. O. 1994. Taxonomic catalog of the ant subfamilies Aneuretinae and Dolichoderinae (Hymeno ...

'' and ''Camponotus'' (''Tanaemyrmex'') sp. The authors suggested that the relatively large-bodied members of the genera '' Camponotus'' and ''

Polyrhachis ''Polyrhachis'' is a genus of formicine ants found in the Old World with over 600 species. The genus is yet to be comprehensively resolved and contains many varied species including nest-weavers (e.g. ''Polyrhachis dives''), swimming workers ( ...

'' may be of particular importance to the plant in terms of nutrient intake. Also among the prey assemblage of ''N. bokorensis'' was the tramp species ''

Cardiocondyla wroughtonii ''Cardiocondyla'' is an Old World genus of ants in the subfamily Myrmicinae. Distribution Approximately 70 species are currently recognized as belonging to this genus, most of which are distributed in the Old World tropics and subtropics, b ...

'' (actually a species complex). The ant-trapping habit of ''N. bokorensis'' is reflected in the local Khmer name for the plant, ''ampuong sramoch'', which means "ants' pithole". This name is not specific to ''N. bokorensis'' but refers to ''Nepenthes'' in general, and is used in

and the town of Pursat. The pitcher fluid of this species is strongly

ic; the label on the herbarium specimen ''Middleton & Monyrak 589'' states that it has a pH of 2.7.

Conservation

Cambodia ranks among the countries with the highest deforestation rates worldwide. Despite this, the habitat of ''N. bokorensis'' remained relatively undisturbed until 2009, lying as it did within the boundaries of

. At the time, François Mey assessed the conservation status of ''N. bokorensis'' as "potentially vulnerable" based on the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

criteria, citing increasing tourism and

land development Land development is the alteration of landscape in any number of ways such as: * Changing landforms from a natural or semi-natural state for a purpose such as agriculture or housing * Subdividing real estate into lots, typically for the purpose ...

as potential threats (Mount Bokor had been leased for private development by the

government of Cambodia The politics of Cambodia are defined within the framework of a constitutional monarchy, in which the king serves as the head of state, and the prime minister is the head of government. The collapse of communism set in motion events that led to th ...

). Habitat loss accelerated rapidly in the following two years as work on the massive "Bokor City" project began in earnest.Mey, F.S. 2011
Works on Phnom Bokor. 2011
''Strange Fruits: A Garden's Chronicle'', August 29, 2011.Mey, F.S. 2011

''Strange Fruits: A Garden's Chronicle'', August 29, 2011. As of 2011, both the type population and the two newly discovered sites are threatened by the ongoing development. Recently, a new threat has been identified. People, particularly women, have taken to picking the plants and having their photos taken with them because while the leaves are still developing, the mouths of the plants resemble men's genitalia. The Cambodian Ministry of Environment has taken to social media to plead with the public to stop picking the plants, which it says could be driven to extinction if people continue to harvest them, as the pitchers allow the plants to obtain nutrients they cannot obtain otherwise.Pester, Patrick, 2022.
Stop picking carnivorous penis plants, Cambodian environmental officials plead
, ''Live Science'', May 17, 2022.

Related species

''Nepenthes bokorensis'' is most closely allied to several other Indochinese pitcher plants, particularly '' N. kampotiana'', '' N. smilesii'', and '' N. thorelii''. It can be distinguished from all three on the basis of its wider, more oblong-shaped lamina and occasional two-flowered partial peduncles. Nepenthes bokorensis lowers

The indumentum of ''N. bokorensis'' is also distinctive, and distinguishes it from ''N. kampotiana'', which typically has glabrous leaves. Compared to ''N. smilesii'', ''N. bokorensis'' has more robust and colourful pitchers with a broader peristome and longer tendrils. The species differs from the enigmatic ''N. thorelii'' in several aspects of vegetative morphology. Firstly, the lamina of ''N. bokorensis'' is

to sub-

and only slightly

down the stem, if at all. In comparison, ''N. thorelii'' has an amplexicaul leaf attachment and the base of the lamina is decurrent into two wings that extend up to 2.5 cm down the stem.Jebb, M.H.P. & M.R. Cheek 1997. A skeletal revision of ''Nepenthes'' (Nepenthaceae). ''Blumea'' 42(1): 1–106. In addition, ''N. thorelii'' has wholly ovoid lower pitchers, whereas those of ''N. bokorensis'' are only

in the basal third, becoming cylindrical above. ''Nepenthes bokorensis'' also appears to be closely related to '' N. kerrii'' of Thailand.Catalano, M. 2010
''Nepenthes kerrii'' M. Catal. et T. Kruetr. sp. nov.
In: '' Nepenthes della Thailandia: Diario di viaggio''. Prague. p. 32. It can be distinguished from this species on the basis of its laminae, which are linear to

as opposed to obovate in the latter. It also differs in having a variable indumentum covering all vegetative and floral parts. In contrast, the indumentum of ''N. kerrii'' is restricted to the leaf axils. The

of ''N. kerrii'' is also considerably shorter than that of ''N. bokorensis''.

Notes

:a.''Geoffray 324'', ''Geoffray 325'' and ''Geoffray 328'' were collected on January 14, 1904, from Popokvil falls, Mount Bokor, at an altitude of 960 m. They are deposited at the

(P) in

. ''Geoffray 324'' consists of upper pitchers with leaves, while ''Geoffray 325'' and ''Geoffray 328'' include lower pitchers with leaves. :b.''Vidal 4780'' was collected on December 20, 1965, at an unknown altitude on Mount Bokor. It consists of a plant with upper pitchers and is deposited at the Museum National d'Histoire Naturelle (P). :c.''M. Martin 1231bis'' was collected on December 7, 1968, on Mount Bokor at an altitude of 800 m. It includes female floral material and lower pitchers, and is deposited at the Museum National d'Histoire Naturelle (P). It is the designated holotype of ''N. bokorensis''. :d.''Middleton & Monyrak 589'' was collected on March 7, 2001, on Mount Bokor at an altitude of 944 m. It consists of two rosette plants and is deposited at the Museum National d'Histoire Naturelle (P). :e.The original Latin description of ''N. bokorensis'' reads:

''N. thorelii'' Lecomte simile, sed foliis longioris latioris oblongis sessilibus vel subpetiolatis basaliter amplexicaulibus peristomio robusto cylindrico pedicellis interdum 2-floribus differt.

:f.''Aug. Chevalier 36411'' was collected on December 15, 1917, from the

of Kampot province. It is deposited at the Museum National d'Histoire Naturelle (P). :g.Three specimens deposited at the Forest Herbarium, Bangkok (BKF) have been tentatively identified as ''N. bokorensis'' from photographs. These are specimens SN 093094, SN 098240, and a third specimen whose number is unknown. The collector of this material is also uncertain.

References

External links

at the Carnivorous Plant Photofinder {{Taxonbar, from=Q9287208 Carnivorous plants of Asia bokorensis Endemic flora of Cambodia Threatened flora of Asia Plants described in 2009