Neomphaloidea is a
superfamily
SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
of deep-
sea snail
Sea snail is a common name for slow-moving marine gastropod molluscs, usually with visible external shells, such as whelk or abalone. They share the taxonomic class Gastropoda with slugs, which are distinguished from snails primarily by the ...
s or
limpet
Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...
s,
marine gastropod
The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda ().
This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is e ...
s. Neomphaloidea is the only superfamily in the order Neomphalida.
The order Neomphalida has the largest ''
in situ
''In situ'' (; often not italicized in English) is a Latin phrase that translates literally to "on site" or "in position." It can mean "locally", "on site", "on the premises", or "in place" to describe where an event takes place and is used in ...
'' radiation in
hydrothermal vent
A hydrothermal vent is a fissure on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hotspot ...
habitats. Neomphalida is a major
taxonomic
Taxonomy is the practice and science of categorization or classification.
A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
grouping of
sea snails
Sea snail is a common name for slow-moving marine gastropod molluscs, usually with visible external shells, such as whelk or abalone. They share the taxonomic class Gastropoda with slugs, which are distinguished from snails primarily by the ...
, vent-endemic
marine gastropod
The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda ().
This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is e ...
s that form a very ancient lineage, going back to the
Palaeozoic era.
2005 taxonomy
The superfamily Neomphaloidea was regarded for a long time as belonging within the clade
Vetigastropoda
Vetigastropoda is a major Taxonomy (biology), taxonomic group of sea snails, marine (ocean), marine gastropod mollusc, mollusks that form a very ancient lineage (evolution), lineage. Taxonomically the Vetigastropoda are sometimes treated as an O ...
. Superfamily Neomphaloidea was also classified in the clade
Vetigastropoda
Vetigastropoda is a major Taxonomy (biology), taxonomic group of sea snails, marine (ocean), marine gastropod mollusc, mollusks that form a very ancient lineage (evolution), lineage. Taxonomically the Vetigastropoda are sometimes treated as an O ...
according to the
taxonomy of the Gastropoda by Bouchet & Rocroi, 2005.
2010 taxonomy
Molecular phylogeny showed however that it belongs in its own order , the Neomphalida, and that this clade is
basal to the Vetigastropoda. The Neomphalina is, based on optimal phylogenetic analysis, a monophyletic clade, with uncertain relations among the gastropods.
Description
The anatomical characters of the Neomphaloidea largely follow the patterns as in the Vetigastropoda. However, unusual morphological and phylogentic characters suggest a different systematic position and place it in its own order, the Neomphalida. The formal placement of Neomphalida within the Gastropoda however remains ambiguous.
[Stephanie W. Aktipis & Gonzalo Giribet (2010). "A phylogeny of Vetigastropoda and other “archaeogastropods”: re-organizing old gastropod clades". ''Invertebrate Biology'' 129(3): 220-240](_blank)
.
Families
Families within the Neomphaloidea include:
*
Melanodrymiidae
Melanodrymiidae is a family of gastropods belonging to the superfamily Neomphaloidea.Bouchet, P. (2011). Melanodrymiidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=411613 on 2012- ...
Salvini-Plawen & Steiner, 1995
*
Neomphalidae
Neomphalidae is a family of sea snails or limpets, specifically deep sea hydrothermal vent limpets. This family was included in the Vetigastropoda, which is a clade according to the Bouchet & Rocroi, 2005. It is now included in the clade Neomp ...
McLean, 1981
*
Peltospiridae
Peltospiridae is a small family of gastropods that used to belong to the clade Vetigastropoda (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005), but is now included in the clade NeomphalinaBouchet, P.; Gofas, S. (2010). Pel ...
McLean, 1989
A few genera within Neomphaloidea have been unassigned to a family:
* ''
Helicrenion'' Warén & Bouchet, 1993
* ''
Retiskenea
''Retiskenea'' is a genus of sea snails, marine gastropod mollusks in the Superfamily (zoology), superfamily Neomphaloidea.Bouchet, P. (2010). Retiskenea Warén & Bouchet, 2001. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollu ...
'' Warén & Bouchet, 2001
[Bouchet, P. (2010). ''Retiskenea'' Warén & Bouchet, 2001. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=449910 on 2017-05-26]
Overview of species
Species within the Neomphaloidea include:
# ''
Leptogyra alaskana
''Leptogyra'' is a genus of sea snails, marine gastropod mollusks in the family Melanodrymiidae, belonging to the clade Neomphalina.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molec ...
'' Bartsch, 1910
# ''
Leptogyra constricta
''Leptogyra'' is a genus of sea snails, marine gastropod mollusks in the family Melanodrymiidae, belonging to the clade Neomphalina.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molec ...
'' B. A. Marshall, 1988
# ''
Leptogyra costellata
''Leptogyra costellata'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molecular phylogeny of ''L ...
'' Warén & Bouchet, 2009
# ''
Leptogyra eritmeta
''Leptogyra eritmeta'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molecular phylogeny of ''Lept ...
'' Bush, 1897
# ''
Leptogyra inconspicua
''Leptogyra inconspicua'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molecular phylogeny of ''L ...
'' Bush, 1897
# ''
Leptogyra inflata
''Leptogyra inflata'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molecular phylogeny of ''Lepto ...
'' Warén & Bouchet, 1993
# ''
Leptogyra patula
''Leptogyra'' is a genus of sea snails, marine gastropod mollusks in the family Melanodrymiidae, belonging to the clade Neomphalina.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molec ...
'' B. A. Marshall, 1988
# ''
Leptogyra verrilli
''Leptogyra verrilli'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and molecular phylogeny of ''Lept ...
'' Bush, 1897
# ''
Leptogyropsis inflata
''Leptogyropsis'' is a genus of sea snails, marine gastropod mollusks in the family Melanodrymiidae, belonging to the clade Neomphalina.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and mol ...
'' Hasegawa, 1997
# ''
Leptogyropsis kalinovoae
''Leptogyropsis'' is a genus of sea snails, marine gastropod mollusks in the family Melanodrymiidae, belonging to the clade Neomphalina.Hess H., Beck F., Gensler H., Kano Y., Kiel S. & Haszprunar G. (2008). "Microanatomy, shell structure and m ...
'' B. A. Marshall, 1988
# ''
Leptogyropsis kaltanae'' B. A. Marshall, 1988
# ''
Melanodrymia aurantiaca
''Melanodrymia aurantiaca'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae
Melanodrymiidae is a family of gastropods belonging to the superfamily Neomphaloidea.Bouchet, P. (2011). Melanodrymiidae. Accesse ...
'' Hickman, 1984
# ''
Melanodrymia brightae
''Melanodrymia brightae'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae
Melanodrymiidae is a family of gastropods belonging to the superfamily Neomphaloidea.Bouchet, P. (2011). Melanodrymiidae. Accessed t ...
'' Warén & Bouchet, 1993
# ''
Melanodrymia galeronae
''Melanodrymia galeronae'' is a species of sea snail, a marine gastropod mollusk in the family Melanodrymiidae.
Description
The length of the shell attains 3 mm. It is spirallic in shape.
Distribution
This marine species occurs on the East P ...
'' Warén & Bouchet, 2001
# ''
Xyleptogyra kapalae'' B. A. Marshall, 1988
# ''
Cyathermia naticoides
''Cyathermia naticoides'' is a species of sea snail, a marine gastropod mollusk in the family Neomphalidae.
Description
The length of the shell attains 6.6 mm.
Distribution
This marine species was found on the East Pacific Rise
The East ...
'' Warén & Bouchet, 1989
# ''
Lacunoides exquisitus
''Lacunoides exquisitus'' is a species of sea snail, a marine gastropod mollusk in the family Neomphalidae
Neomphalidae is a family of sea snails or limpets, specifically deep sea hydrothermal vent limpets. This family was included in the ...
'' Warén & Bouchet, 1989
# ''
Lacunoides vitreus'' Warén & Bouchet, 2001
# ''
Lamellomphalus manusensis'' S.-Q. Zhang & S.-P. Zhang, 2017
# ''
Neomphalus fretterae'' McLean, 1981
# ''
Planorbidella depressa'' Warén & Bouchet, 1993
# ''
Planorbidella planispira'' (Warén & Bouchet, 1989)
# ''
Solutigyra reticulata'' Warén & Bouchet, 1989
# ''
Symmetromphalus hageni
''Symmetromphalus hageni'' is a species of sea snail in the family Neomphalidae.
Distribution
''Symmetromphalus hageni'' lives in hot vent sites in the Bismarck Sea
The Bismarck Sea (, ) lies in the southwestern Pacific Ocean within the n ...
'' L. Beck, 1992
# ''
Symmetromphalus regularis
''Symmetromphalus regularis'' is a species of sea snail, a marine gastropod mollusk in the family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other rela ...
'' McLean, 1990
# ''
Chrysomallon squamiferum
''Chrysomallon squamiferum'', common name, commonly known as the scaly-foot gastropod, scaly-foot snail, sea pangolin, or volcano snail is a species of deep-sea hydrothermal vent, hydrothermal-vent snail, a marine (ocean), marine gastropod mollu ...
'' Chen, Linse, Copley & Rogers, 2015
# ''
Ctenopelta porifera'' Warén & Bouchet, 1993
# ''
Depressigyra globulus'' Warén & Bouchet, 1989
# ''
Echinopelta fistulosa'' McLean, 1989
# ''
Gigantopelta aegis'' Chen, Linse, Roterman, Copley & Rogers, 2015
# ''
Gigantopelta chessoia'' Chen, Linse, Roterman, Copley & Rogers, 2015
# ''
Hirtopelta hirta'' McLean, 1989
# ''
Hirtopelta tufari'' L. Beck, 2002
# ''
Lirapex costellatus'' Warén & Bouchet, 2001
# ''
Lirapex granularis'' Warén & Bouchet, 1989
# ''
Lirapex humatus'' Warén & Bouchet, 1989
# ''
Nodopelta heminoda'' McLean, 1989
# ''
Nodopelta rigneae'' Warén & Bouchet, 2001
# ''
Nodopelta subnoda'' McLean, 1989
# ''
Pachydermia laevis'' Warén & Bouchet, 1989
# ''
Pachydermia sculpta'' Warén & Bouchet, 1993
# ''
Peltospira delicata
''Peltospira delicata'' is a species of sea snail, a marine gastropod mollusk in the family Peltospiridae.
Description
Distribution
This species occurs in the Pacific Ocean off Mexico
Mexico (Spanish language, Spanish: México), offic ...
'' McLean, 1989
# ''
Peltospira lamellifera
''Peltospira lamellifera'' is a species of sea snail, a marine gastropod mollusk in the family Peltospiridae
Peltospiridae is a small family of gastropods that used to belong to the clade Vetigastropoda (according to the taxonomy of the Gast ...
'' Warén & Bouchet, 1989
# ''
Peltospira operculata'' McLean, 1989
# ''
Peltospira smaragdina'' Warén & Bouchet, 2001
# ''
Rhynchopelta concentrica
''Rhynchopelta concentrica'' is a species of sea snail, a marine gastropod mollusk in the family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relat ...
'' McLean, 1989
# ''
Helicrenion reticulatum'' Warén & Bouchet, 1993
# ''
Retiskenea diploura'' Warén & Bouchet, 2001
See also
The other superfamily of hydrothermal vent limpets is the
Lepetodriloidea
Lepetodriloidea is a superfamily of small to large sea snails, hydrothermal vent limpets, marine gastropod mollusks in the clade Vetigastropoda
Vetigastropoda is a major taxonomic group of sea snails, marine gastropod mollusks that form ...
.
References
External links
Steffen Kiel (2010), ''The Vent and Seep Biota: Aspects from Microbes to Ecosystems'', NatureThomas Kunze, Martin Hess, Martin Brückner, Friederike Beck & Gerhard Haszprunar, ''Skeneimorph gastropods in Neomphalina and Vetigastropoda — A preliminary report'', Zoosymposia 1: 119–131 (2008); ISSN 1178-9913
{{Taxonbar, from=Q874910
Gastropod superfamilies
Neomphalina
Marine gastropods