''Neogaeornis'' is a controversial prehistoric

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of diving

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

. The single known species, ''Neogaeornis wetzeli'', was described from

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s found in the

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...

Quiriquina Formation The Quiriquina Formation is a geological formation in Chile whose strata date back to the Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. ...

Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...

. It lived about 70-67

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

. It remains known from the single

tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) and meta ...

described in 1929 by Lambrecht, and today housed in the Paläontologisches Institut und Museum in

Kiel Kiel () is the capital and most populous city in the northern Germany, German state of Schleswig-Holstein, with a population of 246,243 (2021). Kiel lies approximately north of Hamburg. Due to its geographic location in the southeast of the J ...

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

.Chiappe (1991)

Taxonomy

It is controversial because of its uncertain placement. While clearly related to modern birds, ''N. wetzeli'' might not be a particularly close relative, but rather belong to the Baptornithidae, a member of the flightless and toothed

Hesperornithes Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as '' Hesperorn ...

. If this is so, ''Neogaeornis'' is among the very youngest records of this lineage, and the first one from the Southern Hemisphere. However, the Hesperornithiformes are known to have been birds of the open epicontinental and

shelf sea A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...

s which avoided the outer oceans as surrounded South America at that time. And though apparently somewhat migratory, they are only known from

temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...

to warm

subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...

climates, and it seems that towards the end of the Cretaceous their range shifted polewards. Others consider it a close relative of certain modern birds, such as the

Gaviiformes Gaviiformes is an order of aquatic birds containing the loons or divers and their closest extinct relatives. Modern gaviiformes are found in many parts of North America and northern Eurasia (Europe, Asia and debatably Africa), though prehistoric ...

(loons/divers). Both theories are problematic, as neither group is known from the Southern Hemisphere. The even more controversial supposed loon ancestor ''

Polarornis ''Polarornis'' is a genus of prehistoric bird, possibly an anserimorph. It contains a single species ''Polarornis gregorii'', known from incomplete remains of one individual found on Seymour Island, Antarctica, in rocks which are dated to the L ...

'' from

Seymour Island Seymour Island or Marambio Island, is an island in the chain of 16 major islands around the tip of the Graham Land on the Antarctic Peninsula. Graham Land is the closest part of Antarctica to South America. It lies within the section of the isla ...

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...

presents a similar dilemma and ''Neogaeornis'' and there is little consensus about the age of ''Polarornis'', and so all that can be said is that ''Polarornis'' and ''Neogaeornis'' were similarly sized birds with similar lifestyles. ''N. wetzeli'' may also be the ancestor to the

grebe Grebes () are aquatic diving birds in the order Podicipediformes . Grebes are widely distributed freshwater birds, with some species also found in marine habitats during migration and winter. Some flightless species exist as well, most notably ...

s or Podicipediformes. Most recent phylogenetic studies seem to favour its position as a basal

loon Loons (North American English) or divers (British / Irish English) are a group of aquatic birds found in much of North America and northern Eurasia. All living species of loons are members of the genus ''Gavia'', family Gaviidae and order Gavi ...

.Carolina Acosta Hospitaleche, Javier N. Gelfo, New Antarctic findings of Upper Cretaceous and lower Eocene loons (Aves: Gaviiformes), Annales de Paléontologie Volume 101, Issue 4, October–December 2015, Pages 315–324 Alongside ''

'' and some yet unnamed

Antarctic The Antarctic ( or , American English also or ; commonly ) is a polar region around Earth's South Pole, opposite the Arctic region around the North Pole. The Antarctic comprises the continent of Antarctica, the Kerguelen Plateau and other ...

specimens, it seems to suggest a

Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...

n origin for this group. In 2017 a phylogenetic study Agnolín and colleagues have found ''Neogaeornis'' to be stem-

anseriform Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which inc ...

s along with ''

'', ''

Australornis ''Australornis'' (Latin: "southern bird") is a genus of extinct seabird discovered in New Zealand. It lived in the Paleocene epoch, 60.5 to 61.6 million years ago (Ma). The type species name originates from ''australis'', Latin for "southern", a ...

'' and ''

Vegavis ''Vegavis'' is a genus of extinct bird that lived during the Late Cretaceous (Maastrichtian stage) of Antarctica, some 68 to 66 mya. Among modern birds, most studies show that ''Vegavis'' is most closely related to ducks and geese (Anatidae), b ...

'' in the family

Vegaviidae Vegaviidae is an extinct family of ornithurines, often regarded as stem-anseriforms, which existed during the Late Cretaceous and possibly the Paleocene. Fossils attributed to the family have been found in Canada, Chile, New Zealand, and Antarct ...

References

Bibliography

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Taxonomy

References

Bibliography

Further reading