The Negro–Branco moist forests (NT0143) is an
ecoregion
An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...
of tropical moist
broadleaf forest
A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
to the east of the
Andes
The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the List of longest mountain chains on Earth, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range ...
in southern Venezuela, eastern Colombia and northern Brazil, in the
Amazon biome
The Amazon biome () contains the Amazon rainforest, an area of tropical rainforest, and other ecoregions that cover most of the Amazon basin and some adjacent areas to the north and east. The biome contains blackwater river, blackwater and whitewa ...
. It lies on the watershed between the
Orinoco
The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers approximately 1 million km2, with 65% of it in Venezuela and 35% in Colombia. It is the List of rivers by discharge, f ...
and
Rio Negro basins. It includes both
blackwater and
whitewater
Whitewater forms in the context of rapids, in particular, when a river's Stream gradient, gradient changes enough to generate so much turbulence that air is trapped within the water. This forms an unstable current that foam, froths, making t ...
rivers, creating different types of seasonally flooded forest. The vegetation is more typical of the
Guiana
The Guianas, also spelled Guyanas or Guayanas, are a geographical region in north-eastern South America. Strictly, the term refers to the three Guianas: Guyana, Suriname, and French Guiana, formerly British, Dutch, and French Guiana respectiv ...
region than the Amazon.
Location
The ecoregion has an area of
It overlays the Casiquiare Rift.
The center of the ecoregion is in the extreme south of Venezuela, and covers part of the
Parima Tapirapecó National Park
Parima Tapirapecó National Park (''Parque Nacional Parima Tapirapecó'') is a Venezuelan national park in the southern state of Amazonas.
Location
The Parima Tapirapecó National Park is in the Atabapo and Río Negro municipalities. It is ...
.
It extends west into Colombia, where it includes the south of
El Tuparro National Natural Park
El Tuparro National Natural Park () is a national park located in the Vichada Department in the Orinoquía Region of Colombia. It is the only protected area in the Eastern Plains (''Llanos Orientales'') under Colombia's Natural Parks System.
On ...
, and extends to the southeast into the Brazilian states of
Amazonas and
Roraima
Roraima ( ) is one of the 26 states of Brazil. Located in the country's North Region, it is the northernmost and most geographically and logistically isolated state in Brazil. It is bordered by the state of Pará to the southeast, Amazonas t ...
, including part of the
Pico da Neblina National Park
Pico da Neblina National Park () is a national park in the state of Amazonas in the north of Brazil, bordering on Venezuela. It overlaps with several indigenous territories, which creates tensions over land use, as does the military presence due ...
.
It adjoins the
Caquetá and
Japurá–Solimões–Negro moist forests to the south, the
Llanos
The Llanos ( Spanish ''Los Llanos'', "The Plains"; ) is a vast tropical grassland plain situated to the east of the Andes in Colombia and Venezuela, in northwestern South America. It is an ecoregion of the tropical and subtropical grasslands, ...
grasslands to the north, the
Guianan piedmont and lowland moist forests
The Guianan piedmont and lowland moist forests (NT0182) is an ecoregion in the south of Venezuela and the north of Brazil. It is in the Amazon biome.
The ecoregion is relatively intact, largely protected by conservation units or indigenous territ ...
to the east and the
Uatuma–Trombetas moist forests
The Uatuma–Trombetas moist forests (NT0173) is an ecoregion in northwest Brazil in the Amazon biome.
It covers the Amazon basin north of the Amazon River from close to the Atlantic Ocean to the Rio Negro west of Manaus.
The ecoregion is relati ...
to the southeast.
It also adjoins or contains areas of
Rio Negro campinarana
Campinarana (NT0158, ), also called Rio Negro Campinarana, is a neotropical ecoregion in the Amazon biome of the north west of Brazil, southern Venezuela, and the east of Colombia that contains vegetation adapted to extremely poor soil. It include ...
.
Hydrology
The Negro-Branco moist forests ecoregion is on the watershed between the
Orinoco
The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers approximately 1 million km2, with 65% of it in Venezuela and 35% in Colombia. It is the List of rivers by discharge, f ...
in Venezuela and the
Rio Negro, which is known as the
Guainía River in Colombia and is a major tributary of the
Amazon River.
Rivers include nutrient-poor
clearwater and
blackwater rivers, the latter type stained dark by tannin, and nutrient-rich
whitewater
Whitewater forms in the context of rapids, in particular, when a river's Stream gradient, gradient changes enough to generate so much turbulence that air is trapped within the water. This forms an unstable current that foam, froths, making t ...
rivers.
The blackwater
Vichada River
The Vichada River (, ) is a blackwater river in Colombia, South America. It flows into the Orinoco River.
The eastward course of the Vichada is offset by an Impact crater, impact structure, called the Vichada Structure. The structure is most li ...
forms the northern border in Colombia, and the clearwater
Ventuari River
The Ventuari River is the largest tributary of the Orinoco in southern Venezuela. The Ventuari flows from south-central Venezuela in the Guiana Highlands southwest into the Orinoco River. It is long and its major tributary is the Manapiare River ...
forms the northern border in Venezuela, both tributaries of the
Orinoco
The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers approximately 1 million km2, with 65% of it in Venezuela and 35% in Colombia. It is the List of rivers by discharge, f ...
.
In Colombia the whitewater
Guaviare and
Inírida rivers are in the Orinoco Basin.
The southern border in Colombia is defined by the middle Guaviare and by the upper reaches of the blackwater Guainía / Rio Negro to the Venezuela-Colombia border.
In Venezuela the blackwater
Casiquiare canal
The Casiquiare river or canal () is a natural distributary of the upper Orinoco flowing southward into the Rio Negro, in Venezuela, South America. As such, it forms a unique natural canal between the Orinoco and Amazon river systems. It is the ...
, a
distributary
A distributary, or a distributary channel is a stream channel that branches off and flows a main stream channel. It is the opposite of a ''tributary'', a stream that flows another stream or river. Distributaries are a result of river bifurc ...
of the Orinoco, feeds the Guainia/Negro River.
The Rio Negro defines the southern border in Brazil to the whitewater
Rio Branco.
Environment
The ecoregion is on the ancient
Guiana Shield
The Guiana Shield (; ; ; ) is one of the three cratons of the South American Plate. It is a 1.7 billion-year-old Precambrian geological formation in northeast South America that forms a portion of the northern coast. The higher elevations on ...
, a
craton
A craton ( , , or ; from "strength") is an old and stable part of the continental lithosphere, which consists of Earth's two topmost layers, the crust and the uppermost mantle. Having often survived cycles of merging and rifting of contine ...
formed in the
Precambrian
The Precambrian ( ; or pre-Cambrian, sometimes abbreviated pC, or Cryptozoic) is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of t ...
.
Elevations range from in the west to over in the east in Venezuela.
Terrain includes lowland plains, rolling hills and low sandstone plateaus.
Until recently the lowland plains of the region were covered in lakes and seas, which deposited layers of sediment.
Soil types are generally poor in nutrients, mainly
podzol
Podzols, also known as podosols, spodosols, or espodossolos, are the typical soils of coniferous or boreal forests and also the typical soils of eucalypt forests and heathlands in southern Australia. In Western Europe, podzols develop on heathlan ...
s on the old alluvial terraces.
The ecoregion is part of the Río Negro-Juruá Moist Forests, a global ecoregion, the other parts being the Caquetá,
Solimões–Japurá and Japurá–Solimoes–Negro moist forests.
The reasonably intact global ecoregion has high annual rainfall, diverse soils and varied terrain, resulting in a high level of biodiversity.
It has not been studied in great depth by scientists.
Flora

The Negro–Branco moist forests are in the
tropical and subtropical moist broadleaf forests
Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF).
Description
TSMF is generally found in la ...
biome of the
neotropical realm
The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropics, tropical Ecoregion#Terrestrial, terrestrial ecoregions of the Americas and the entire South American temperat ...
.
Vegetation includes flooded and terrra firme lowland moist forest, tall or medium-tall
campinarana
Campinarana (NT0158, ), also called Rio Negro Campinarana, is a neotropical ecoregion in the Amazon biome of the north west of Brazil, southern Venezuela, and the east of Colombia that contains vegetation adapted to extremely poor soil. It includ ...
lowland forest, patches of shrubs and herbaceous
savanna
A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
-like meadows.
Forests are typical of the Guiana region, with a different mix of plants from the classical
Amazon rainforest
The Amazon rainforest, also called the Amazon jungle or Amazonia, is a Tropical and subtropical moist broadleaf forests, moist broadleaf tropical rainforest in the Amazon biome that covers most of the Amazon basin of South America. This basin ...
.
Humiriaceae
Humiriaceae (or, alternatively Houmiriaceae Juss.) is a family of evergreen flowering plants. It comprises 8 genera and 56 known species. The family is exclusively Neotropical, except one species found in tropical West Africa
West Africa, al ...
,
Rapateaceae
The Rapateaceae are a family of flowering plants. The botanical name has been recognized by most taxonomists.
The APG II system of 2003 also recognizes this family, and assigns it to the order Poales in the clade commelinids, in the monocots. T ...
,
Tepuianthaceae,
Theaceae
Theaceae (), the tea family, is a family of flowering plants comprising shrubs and trees, including the economically important tea plant, and the ornamental camellias. It can be described as having from seven to 40 genera, depending on the sour ...
and
Xyridaceae
The Xyridaceae are a family of flowering plants. This family has been recognized by many taxonomists and is known as the yellow-eyed grass family.
The APG II system, of 2003 (unchanged from the APG system of 1998), also recognizes this family, ...
are common families that do not belong to the Amazon flora.
The forests hold relatively few
epiphyte
An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
s or
liana
A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...
s compared to other parts of the western Amazon region.
The flat Casiquiare peneplain in Venezuela holds forests, savannas and other formations.
It contains blackwater and whitewater rivers, which create
igapó
(, from Tupi language, Old Tupi: "root forest") is a word used in Brazil for Blackwater river, blackwater-flooded forests in the Amazon biome. These forests and similar swamp forests are seasonally inundated with freshwater. They typically occur ...
and
várzea forest A várzea forest is a seasonal floodplain forest inundated by whitewater rivers that occurs in the Amazon biome. Until the late 1970s, the definition was less clear and várzea was often used for all periodically flooded Amazonian forests.
Althoug ...
along their flooded banks.
There are seasonally flooded and terra firme evergreen lowland forests reaching , and low evergreen flooded palm forests that reach .
The dominant trees of the low palm forests are ''
Mauritia flexuosa
''Mauritia flexuosa'', known as the moriche palm, ''ité'' palm, ''ita'', ''buriti'', ''muriti'', ''miriti'' (Brazil), ''canangucho'' (Colombia), ''morete or acho'' (Ecuador), ''palma real'' (Bolivia), or ''aguaje'' (Peru), is a Arecaceae, palm t ...
,
Mauritiella aculeata'', and dense groups of ''
Euterpe catinga
''Euterpe catinga'' is a palm species in the genus ''Euterpe''. It is found in forests of a dry, sandy soil and very peculiar vegetation, known as catinga forests or Campinarana in northern South America (Guyana, Venezuela, Colombia, Ecuador, Per ...
,
Iriartea setigera'', and ''
Socratea exorrhiza
''Socratea exorrhiza'', the walking palm or cashapona, is a palm native to rainforests in tropical Central and South America. It can grow to 25 metres in height, with a stem diameter of up to 16 cm, but is more typically 15–20 m tall ...
''.
The terra firme forest holds ''
Lecointea amazonica
''Lecointea'' is a genus of flowering plants in the family Fabaceae. It contains seven species native to the tropical Americas.
* ''Lecointea amazonica'' Ducke
* ''Lecointea guianensis''
* ''Lecointea hatschbachii''
* ''Lecointea lasiogyne''
* ...
,
Clathrotropis glaucophylla,
Peltogyne venosa'', and species in the ''
Ocotea
''Ocotea'' is a genus of flowering plants belonging to the family Lauraceae. Many are evergreen trees with lauroid leaves.
There are over 520 species currently accepted within the genus, distributed mostly in tropical and subtropical areas of ...
,
Nectandra
''Nectandra'' is a genus of plant in the family Lauraceae. They are primarily Neotropical, with ''Nectandra coriacea'' being the only species reaching the southernmost United States. They have fruit with various medical effects. Sweetwood is a co ...
,
Licania
''Licania'' is a genus of over 100 species of trees and shrubs in the family Chrysobalanaceae. Species are found naturally occurring in neotropical forests from southern Mexico to Brazil and the Lesser Antilles. Due to increased deforestation an ...
,
Trichilia
''Trichilia'' is a flowering plant genus in the family (biology), family Meliaceae. These plants are particularly diverse in sub-Saharan Africa and tropical South America.
Several species are used in folk medicine and shamanism – e.g. ''Tr ...
,
Guarea
''Guarea'' is a genus of evergreen trees or shrubs in the family Meliaceae, native to tropical Africa and Central and South America. At their largest, they are large trees 20–45 m tall, with a trunk over 1 m diameter, often buttressed at the b ...
,
Toulicia,
Erisma
''Erisma'' is a genus of flowering plants belonging to the family Vochysiaceae
Vochysiaceae is a plant family belonging to the order Myrtales.
Description
Trees or shrubs with opposite leaves; flowers are zygomorph 1-(3)-5 merous; ovary in ...
'', and ''
Ruizterania
''Ruizterania'' is a genus of flowering plants belonging to the family Vochysiaceae
Vochysiaceae is a plant family belonging to the order Myrtales.
Description
Trees or shrubs with opposite leaves; flowers are zygomorph 1-(3)-5 merous; ova ...
'' genera.
The tall canopy forests also hold trees of the ''
Oenocarpus
''Oenocarpus'' is a genus of pinnate-leaved palms (Arecaceae) native to Trinidad, southern Central and tropical South America. (2004): World Checklist of Arecaceae &ndash''Oenocarpus'' The Board of Trustees of the Royal Botanic Gardens, Kew. R ...
,
Socratea
''Socratea'' is a genus of five species of palms found in tropical
The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where ...
,
Leopoldinia'', and ''
Bactris
''Bactris'' is a genus of spiny palms which are native to Mexico, South and Central America and the Caribbean. Most species are small trees about tall, but some are large trees while others are shrubs with subterranean stems. They have simple o ...
'' genera.
There are patches of campinarana on white sands near the border of Venezuela and Colombia.
There is a high level of endemism of flora including the genera ''
Duckeanthus
''Duckeanthus'' is a genus of plant in the family Annonaceae. It is native to Brazil. It contains a single species, ''Duckeanthus grandiflorus''. Robert Elias Fries, the Swedish botanist who first formally described it, named it in honor of Ad ...
,
Heteropetalum,
Pseudephedranthus,
Urospathella,
Aquiaria,
Angostylis,
Astrococcus'', and ''
Chonocentrum
''Chonocentrum'' is a genus of the family Phyllanthaceae
Phyllanthaceae is a Family (biology), family of flowering plants in the eudicot Order (biology), order Malpighiales. It is most closely related to the family Picrodendraceae.Kenneth J. ...
''.
The economically valuable palm ''
Leopoldinia piassaba'' is endemic to the ecoregion.
In the Rio Negro basin the many blackwater streams and small rivers flood a large area of
igapó
(, from Tupi language, Old Tupi: "root forest") is a word used in Brazil for Blackwater river, blackwater-flooded forests in the Amazon biome. These forests and similar swamp forests are seasonally inundated with freshwater. They typically occur ...
forest for 5–6 months each year.
The canopy is up to high.
Common species are ''
Virola elongata,
Eschweilera longipes,
Eschweilera pachysepala,
Aldina latifolia'', and ''
Pithecellobium amplissimum''.
Fauna
There are 194 species of mammals, including a few endemic species such as
golden-backed uakari (''Cacajao melanocephalus''),
black bearded saki (''Chiropotes satanas''),
Tschudi's slender opossum (''Marmosops impavidus''),
least big-eared bat (''Neonycteris pusilla''),
Guianan spear-nosed bat (''Phyllostomus latifolius''),
Eldorado broad-nosed bat (''Platyrrhinus aurarius''),
Venezuelan fish-eating rat (''Neusticomys venezuelae''),
MacConnell's climbing mouse (''Rhipidomys macconnelli''), the guinea pig ''
Cavia guianae
''Cavia guianae'' is a guinea pig species from South America. It is found in southern Venezuela, Guyana, and portions of northern Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is th ...
'' and
Simon's spiny rat (''Proechimys simonsi'').
Common species include
South American tapir
The South American tapir (''Tapirus terrestris''), also commonly called the Brazilian tapir (from the Tupi ), the Amazonian tapir, the maned tapir, the lowland tapir, (Brazilian Portuguese), and ''la sachavaca'' (literally "bushcow", in mixed ...
(''Tapirus terrestris''),
collared peccary
The collared peccary (''Dicotyles tajacu'') is a peccary, a species of artiodactyl (even-toed) mammal in the family Peccary, Tayassuidae found in North America, North, Central America, Central, and South America. It is the only member of the gen ...
(''Pecari tajacu''),
pale-throated sloth
The pale-throated sloth (''Bradypus tridactylus''), occasionally known as the ai (), is a species of three-toed sloth that inhabits tropical rainforests in northern South America.
It is similar in appearance to, and often confused with, the brow ...
(''Bradypus tridactylus''),
long-tailed weasel
The long-tailed weasel (''Neogale frenata''), also known as the bridled weasel, masked ermine, or big stoat, is a species of weasel found in North America, North, Central America, Central, and South America. It is distinct from the Stoat, short-t ...
(''Mustela frenata''),
short-eared dog
The short-eared dog (''Atelocynus microtis''), also known as the small-eared dog, is a unique and elusive canid species endemic to the Amazonian basin. This is the only species assigned to the genus ''Atelocynus''.
Other names
The short-eared ...
(''Atelocynus microtis''),
bush dog
The bush dog (''Speothos venaticus'') is a canine found in Central and South America. In spite of its extensive range, it is very rare in most areas except in Suriname, Guyana and Peru; it was first described by Peter Wilhelm Lund from fossils ...
(''Speothos venaticus'') and three species of small cats of the
Leopardus
''Leopardus'' is a genus comprising eight species of Felinae, small cats native to the Americas. This genus is considered the oldest branch of a genetic lineage of small cats in the Americas whose common ancestor crossed the Bering land bridge fr ...
genus.
There are many reptiles and amphibians.
Snakes include fer-de-lance (''
Bothrops asper
The terciopelo (''Bothrops asper'') is a species of pit vipers, found from north-east Mexico to northern South America. At low to moderate elevations, its habitat ranges from northeastern Mexico to Colombian and Ecuadorian Andes, as well as Venez ...
''), palm pit-vipers (
Bothriechis
''Bothriechis'' is a genus of pit vipers, commonly called palm vipersJohn M. Mehrtens, Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . or palm-pit vipersCampbell JA, species:William W. Lamar, ...
species), coral snakes (
Micrurus
''Micrurus'' is a genus of venomous coral snakes of the family Elapidae.
Geographic range
Species in the genus ''Micrurus'' are endemic to the Americas.
Species
The following 82 species are recognized as being valid.
*'' Micrurus albicinctus' ...
genus), boa constrictors (
Boa constrictor
The boa constrictor (scientific name also ''Boa constrictor''), also known as the common boa, is a species of large, non-venomous, heavy-bodied snake that is frequently kept and bred in captivity. The boa constrictor is a member of the Family (b ...
) and bushmasters (''
Lachesis muta
''Lachesis muta'', also known as the Southern American bushmaster or Atlantic bushmaster, is a pit viper species found in South America, as well as the island of Trinidad in the Caribbean. Two subspecies are currently recognized, including the no ...
'').
There are many
green iguana
The green iguana (''Iguana iguana''), also known as the American iguana or the common green iguana, is a large, arboreal, mostly Herbivory, herbivorous species of lizard of the genus ''Iguana''. Usually, this animal is simply called the iguana. ...
s (''Iguana iguana'') and tegus lizards (
Tupinambis
''Tupinambis'' is a lizard genus which belongs to the Family (biology), family Teiidae and contains eight described species. These large lizards are commonly referred to as tegus (''teiús'' in Portuguese language, Portuguese). ''Tupinambis meria ...
genus).
486 species of birds have been recorded, a relatively low number for the Amazon region.
They include the endemic
grey-legged tinamou (''Crypturellus duidae''),
crestless curassow
The crestless curassow (''Mitu tomentosum'') is a species of bird in the family Cracidae, the chachalacas, guans, and curassows. It is found in Brazil, Colombia, Guyana, and Venezuela.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramun ...
(''Mitu tomentosum''),
double-striped thick-knee
The double-striped thick-knee (''Hesperoburhinus bistriatus'') is a stone-curlew, a group of waders in the family Burhinidae. The vernacular name refers to the prominent joints in the long greenish-grey legs, and ''bistriatus'' to the two stripe ...
(''Burhinus bistriatus''),
oilbird
The oilbird (''Steatornis caripensis''), locally known as the , is a bird species found in the northern areas of South America including the Caribbean island of Trinidad. It is the only living species in the genus ''Steatornis'', the family Stea ...
(''Steatornis caripensis''),
tawny-tufted toucanet
The tawny-tufted toucanet (''Selenidera nattereri'') is a near-passerine bird in the toucan family Ramphastidae. It is found in Brazil, Colombia, Venezuela, and possible Guyana.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Ja ...
(''Selenidera nattereri''),
Orinoco piculet (''Picumnus pumilus''),
Yapacana antbird (''Myrmeciza disjuncta''),
gray-bellied antbird (''Myrmeciza pelzelni''),
spot-tailed nightjar (''Hydropsalis maculicaudus''),
azure-naped jay (''Cyanocorax heilprini''), and
white-naped seedeater (''Dolospingus fringilloides'').
Threats
The ecoregion covers most of the
Alto Orinoco-Casiquiare Biosphere Reserve.
A small part of the ecoregion is in the
Pico da Neblina National Park
Pico da Neblina National Park () is a national park in the state of Amazonas in the north of Brazil, bordering on Venezuela. It overlaps with several indigenous territories, which creates tensions over land use, as does the military presence due ...
.
There are few threats to the ecoregion, which is inaccessible and has no roads.
The forest is largely intact.
People in riverine communities engage in small-scale rotation agriculture.
There is some low-level logging and in some areas the understory is routinely burned by
Brazil nut
The Brazil nut (''Bertholletia excelsa'') is a South American tree in the family Lecythidaceae, and it is also the name of the tree's commercially harvested edible seeds. It is one of the largest and longest-lived trees in the Amazon rainforest. ...
collectors.
Leaves of the ''
Leopoldinia piassaba'' palm are harvested to make brooms for sale internationally, and this may be unsustainable.
References
Bibliography
*
*
*
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{{DEFAULTSORT:Negro-Branco moist forests
Amazon biome
Amazon rainforest
Upper Amazon
Ecoregions of Brazil
Ecoregions of Colombia
Ecoregions of Venezuela
Neotropical tropical and subtropical moist broadleaf forests