Narkidae
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Members of the family Narkidae are commonly known as sleeper rays. They are restricted to the temperate and tropical
Indo-West Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
from
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to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
to
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, and are exclusively
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
and are absent from
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
habitats. They occur from the
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species o ...
to the
continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
and the upper
continental slope A continental margin is the outer edge of continental crust abutting oceanic crust under coastal waters. It is one of the three major zones of the ocean floor, the other two being deep-ocean basins and mid-ocean ridges. The continental margin ...
to a depth of 350 meters, favoring soft-bottomed habitats.


Description

Adult sleeper rays range in size from 8 to 46 cm in length and have flattened oval, circular, or pear-shaped
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
discs. They have naked skin, without
dermal denticle A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as we ...
s or thorns. The snout is moderately elongate and broadly rounded, with the rostral cartilage reduced to a slender medial rod. This distinguishes the narkids from the family Narcinidae, which have somewhat longer snouts supported by broad rostral cartilages. The mouth is straight, very narrow, and not highly protrusible, with strong labial folds and a weak groove around the periphery. The
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
s are placed just forward of the mouth and are connected to it by a broad nasoral groove; the nasal flaps are long and fused into a central nasal curtain that overlaps the mouth. There are a pair of large
kidney The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; blood ...
-shaped electric organs at the base of the pectoral fins, visible through the skin. The genera ''Heteronarce'' and ''Electrolux'' have two large
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
s, ''Narke'', ''Crassinarke'', and ''Typhlonarke'' have a single dorsal fin, and ''Temera'' has none. No other group of electric rays have genera with a variable number of dorsal fins. The tail is moderately long, with a large, rounded, symmetrical
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
. The color is usually brown or reddish brown above, white or brownish below. There may be spots, blotches, or bands on the dorsal surface, but most species lack complex color patterns or
ocelli A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-l ...
.


Biology and ecology

Sleeper rays are bottom-dwelling, slow-moving fishes often found on soft mud or sand. From those species that have been examined, their diet may be restricted to small, soft-bodied, bottom
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s, such as
polychaete worm Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are mad ...
s. They can generate a strong
electric shock Electrical injury is a physiological reaction caused by electric current passing through the body. The injury depends on the density of the current, tissue resistance and duration of contact. Very small currents may be imperceptible or produce ...
as a defense mechanism. Reproduction is
ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
, with
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
hatching inside the mother.Compagno, L.J.V. and Last, P.R. (1999). Narkidae. Sleeper rays. p. 1443-1446. In: K.E. Carpenter and V.H. Niem (eds.) ''FAO identification guide for fishery purposes. The living marine resources of the Western Central Pacific.'' Rome:
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
.


Species

* Genus '' Crassinarke''
Takagi is a Japanese surname meaning "tall tree". Notable people with the surname include: *Akimitsu Takagi (1920–1995), Japanese crime fiction writer *Jon Mamoru Takagi (1942–1984), American martial arts instructor *, Japanese footballer * Hinako T ...
, 1951
** '' Crassinarke dormitor''
Takagi is a Japanese surname meaning "tall tree". Notable people with the surname include: *Akimitsu Takagi (1920–1995), Japanese crime fiction writer *Jon Mamoru Takagi (1942–1984), American martial arts instructor *, Japanese footballer * Hinako T ...
, 1951
(Sleeper torpedo) * Genus ''
Electrolux Electrolux AB () is a Swedish multinational home appliance manufacturer, headquartered in Stockholm. It is consistently ranked the world's second largest appliance maker by units sold, after Whirlpool. Electrolux products sell under a variety ...
'' Compagno & Heemstra, 2007 ** ''
Electrolux addisoni The ornate sleeper-ray (''Electrolux addisoni'') is a species of electric ray in the family Narkidae, and the only member of the genus ''Electrolux''. It lives on reefs feeding on polychaete worms and small crustaceans, it has only been seen by d ...
'' Compagno & Heemstra, 2007 (Ornate sleeper-ray) * Genus '' Heteronarce''
Regan The family name Regan, along with its cognates O'Regan, O Regan, Reagan, and O'Reagan, is an Anglicized form of the Irish surname Ó Riagáin or Ó Ríogáin, from Ua Riagáin. The meaning is likely to have originated in ancient Gaelic ''ri'' ...
, 1921
** '' Heteronarce bentuviai'' ( Baranes & Randall, 1989) (Elat electric ray) ** '' Heteronarce garmani''
Regan The family name Regan, along with its cognates O'Regan, O Regan, Reagan, and O'Reagan, is an Anglicized form of the Irish surname Ó Riagáin or Ó Ríogáin, from Ua Riagáin. The meaning is likely to have originated in ancient Gaelic ''ri'' ...
, 1921
(Natal electric ray) ** '' Heteronarce mollis'' ( Lloyd, 1907) (Soft electric ray) ** '' Heteronarce prabhui''
Talwar The talwar (), also spelled ''talwaar'' and ''tulwar'', is a type of curved sword or sabre from the Indian subcontinent. Etymology and classification The word ''talwar'' originated from the Sanskrit word ''taravāri'' ( sa, तरवारि) ...
, 1981
(Quilon electric ray) * Genus '' Narke'' Kaup, 1826 ** '' Narke capensis'' (
Gmelin Gmelin may refer to: * Gmelin's test, a chemical test * Gmelin database, a German handbook/encyclopedia of inorganic compounds initiated by Leopold Gmelin People * Carl Christian Gmelin (1762–1837), German botanist, author of ''Flora Badensis ...
, 1789)
(Onefin electric ray) ** ''
Narke dipterygia The numbray or spottail sleeper ray (''Narke dipterygia'') is a species of electric ray in the family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affin ...
'' (
Bloch Bloch is a surname of German origin. Notable people with this surname include: A–F * (1859-1914), French rabbi *Adele Bloch-Bauer (1881-1925), Austrian entrepreneur *Albert Bloch (1882–1961), American painter * (born 1972), German motor journal ...
&
Schneider Schneider may refer to: Hospital * Schneider Children's Medical Center of Israel People * Schneider (surname) Companies and organizations * G. Schneider & Sohn, a Bavarian brewery company * Schneider Rundfunkwerke AG, the former owner of th ...
, 1801)
(Numbray) ** '' Narke japonica'' (
Temminck Coenraad Jacob Temminck (; 31 March 1778 – 30 January 1858) was a Dutch aristocrat, zoologist and museum director. Biography Coenraad Jacob Temminck was born on 31 March 1778 in Amsterdam in the Dutch Republic. From his father, Jacob Temminck ...
&
Schlegel Schlegel is a German occupational surname. Notable people with the surname include: * Anthony Schlegel (born 1981), former American football linebacker * August Wilhelm Schlegel (1767–1845), German poet, older brother of Friedrich * Brad Schlege ...
, 1850)
(Japanese sleeper ray) * Genus '' Temera''
J. E. Gray John Edward Gray, Fellow of the Royal Society, FRS (12 February 1800 – 7 March 1875) was a British zoology, zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ...
, 1831
** '' Temera hardwickii''
J. E. Gray John Edward Gray, Fellow of the Royal Society, FRS (12 February 1800 – 7 March 1875) was a British zoology, zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ...
, 1831
(Finless sleeper ray) * Genus '' Typhlonarke'' Waite, 1909 ** '' Typhlonarke aysoni'' ( A. Hamilton, 1902) (Blind electric ray) ** '' Typhlonarke tarakea''
Phillipps Phillipps is both a given name and an English surname. Notable people with the name include: "Phillipps" has also been a shortened version of Philippson, a German surname especially prevalent amongst German Jews and Dutch Jews. People with the g ...
, 1929
(Oval electric ray)


References

* {{Taxonbar, from=Q2245189 Torpediniformes Ray families Taxa named by Henry Weed Fowler