Megalomyrmex
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''Megalomyrmex'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Myrmicinae Myrmicinae is a subfamily of ants, with about 140 extant genera; their distribution is cosmopolitan. The pupae lack cocoons. Some species retain a functional sting. The petioles of Myrmicinae consist of two nodes. The nests are permanent and ...
. The genus is known only from the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropics, tropical Ecoregion#Terrestrial, terrestrial ecoregions of the Americas and the entire South American temperat ...
, where some of the species are specialized parasites or predators of Attini.


Description

It is difficult to characterize morphologically. Bolton (2003) placed it in the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
Solenopsidini Solenopsidini (meaning "pipe-faced") is a tribe of myrmicine ants with about 20 genera. Genera *'' Adelomyrmex'' *''Anillomyrma'' *'' Austromorium'' *'' Baracidris'' *'' Bariamyrma'' *'' Bondroitia'' *'' Cryptomyrmex'' *'' Dolopomyrmex'' ...
, but with multiple exceptions to the diagnostic characters for the tribe. The tribal characters include a bicarinate clypeus and a median clypeal seta. Most ''Megalomyrmex'' species have a smoothly convex clypeus with no trace of the bicarinate condition, and most have abundant clypeal
setae In biology, setae (singular seta ; from the Latin word for "bristle") are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Annelid setae are stiff bristles present on the body. Th ...
with no distinct or differentiated median seta. In Bolton's (1994) key to genera, ''Megalomyrmex'' keys in multiple places because of variability in mandibular dentition. Nevertheless, the genus has a distinctive habitus: the antenna is 12-segmented with a 3-segmented club; the general integument is smooth and shiny without coarse sculpture or dull areas; the promesonotum is evenly arched, without promesonotal groove; the propodeum is usually smoothly curved between dorsal and posterior faces, at most with blunt, broad-based tubercles, and never with spines; and the hind tibial spur is simple. In short, the workers look like a '' Solenopsis'' with ''
Pheidole ''Pheidole'' is a genus of ants that belongs to the ant subfamily Myrmicinae. The genus is widespread and ecologically dominant. It probably includes more than 1000 species. The genus first evolved in the Americas, eventually spreading across th ...
'' antennae. The mandibular dentition varies from a simple set of 5 similar teeth on the masticatory margin, gradually diminishing in size basally, to a condition with 2 large apical teeth followed by up to 12 small denticles. Some species have unusual
alkaloid Alkaloids are a class of basic, naturally occurring organic compounds that contain at least one nitrogen atom. This group also includes some related compounds with neutral and even weakly acidic properties. Some synthetic compounds of similar ...
s. The hypothesis that alkaloids act at a distance converges with what is known for the genera '' Solenopsis'', ''
Monomorium ''Monomorium'' is a genus of ants in the subfamily Myrmicinae. As of 2013 it contains about 396 species. It is distributed around the world, with many species native to the Old World tropics. It is considered to be "one of the more important grou ...
'' and ''Megalomyrmex'' when workers release volatile venom alkaloids by waving their stingers (i.e., gaster flagging) during interspecific encounters causing their enemies to flee. It is close to ''
Monomorium ''Monomorium'' is a genus of ants in the subfamily Myrmicinae. As of 2013 it contains about 396 species. It is distributed around the world, with many species native to the Old World tropics. It is considered to be "one of the more important grou ...
'' and should possibly be included in the latter.


Distribution and habitat

Although widespread in the Neotropics, from southern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
to northern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, ''Megalomyrmex'' species are never abundant. They occur in low to middle elevation wet to dry forest habitats. Some species are free-living with large diffuse nests in the soil (e.g. ''
Megalomyrmex modestus ''Megalomyrmex modestus'' is a Neotropical species of ants in the subfamily Myrmicinae. Known from Costa Rica, Panama, Colombia, Venezuela. In Costa Rica, ''M. modestus'' is found in mature wet forest of the Atlantic slope. Although it occurs a ...
'') or small nests in dead wood (e.g. '' Megalomyrmex drifti''). Others are specialized social parasites or predators of Attini (e.g. ''
Megalomyrmex adamsae ''Megalomyrmex adamsae'' is a Neotropical species of ants in the subfamily Myrmicinae. ''Megalomyrmex adamsae'', known from Panama, is very similar to '' M. symmetochus'', known from Costa Rica to Panama. Biology ''Megalomyrmex adamsae'' is a s ...
'', ''
Megalomyrmex mondabora ''Megalomyrmex mondabora'' is a Neotropical species of ants in the subfamily Myrmicinae. Known from Costa Rica. In Costa Rica this species occurs in wet forest habitats, typically in mature rainforest. Collections are from sea level to 800 m ...
'', ''
Megalomyrmex symmetochus ''Megalomyrmex symmetochus'' is a species of ant in the subfamily Myrmicinae. It is native to Panama. ''M. symmetochus'' was discovered by William M. Wheeler in late July 1924 in the fungus gardens of the attine '' Sericomyrmex amabilis'' of B ...
'', ''
Megalomyrmex wettereri ''Megalomyrmex wettereri'' is a Neotropical species of ants in the subfamily Myrmicinae. ''Megalomyrmex wettereri'' is known from two lowland rainforest sites: Barro Colorado Island in Panama and La Selva Biological Station in Costa Rica. Nest ...
'').


Species

*''
Megalomyrmex acauna ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Brandão, 1990 *''
Megalomyrmex adamsae ''Megalomyrmex adamsae'' is a Neotropical species of ants in the subfamily Myrmicinae. ''Megalomyrmex adamsae'', known from Panama, is very similar to '' M. symmetochus'', known from Costa Rica to Panama. Biology ''Megalomyrmex adamsae'' is a s ...
'' Longino, 2010 *''
Megalomyrmex ayri ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Brandão, 1990 *''
Megalomyrmex balzani ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Emery, 1894 *''
Megalomyrmex bidentatus ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Fernández & Baena, 1997 *''
Megalomyrmex bituberculatus ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' (Fabricius, 1798) *''
Megalomyrmex brandaoi ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Boudinot, Sumnicht & Adams, 2013 *''
Megalomyrmex caete ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Brandão, 1990 *''
Megalomyrmex cuatiara ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Brandão, 1990 *'' Megalomyrmex cupecuara'' Brandão, 1990 *''
Megalomyrmex cyendyra ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphological ...
'' Brandão, 1990 *'' Megalomyrmex drifti'' Kempf, 1961 *''
Megalomyrmex emeryi ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphological ...
'' Forel, 1904 *''
Megalomyrmex foreli ''Megalomyrmex foreli'' (named after Auguste Forel) is a Neotropical species of ants in the subfamily Myrmicinae. Habitat and distribution The species is known from Costa Rica south to Andean regions of Colombia, Ecuador, and northern Peru. ''M ...
'' Emery, 1890 *'' Megalomyrmex fungiraptor'' Boudinot, Sumnicht & Adams, 2013 *'' Megalomyrmex glaesarius'' Kempf, 1970 *'' Megalomyrmex gnomus'' Kempf, 1970 *'' Megalomyrmex goeldii'' Forel, 1912 *'' Megalomyrmex iheringi'' Forel, 1911 *'' Megalomyrmex incisus'' Smith, 1947 *''
Megalomyrmex leoninus ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Forel, 1885 *'' Megalomyrmex longinoi'' Boudinot, Sumnicht & Adams, 2013 *'' Megalomyrmex megadrifti'' Boudinot, Sumnicht & Adams, 2013 *'' Megalomyrmex milenae'' Boudinot, Sumnicht & Adams, 2013 *'' Megalomyrmex miri'' Brandão, 1990 *''
Megalomyrmex modestus ''Megalomyrmex modestus'' is a Neotropical species of ants in the subfamily Myrmicinae. Known from Costa Rica, Panama, Colombia, Venezuela. In Costa Rica, ''M. modestus'' is found in mature wet forest of the Atlantic slope. Although it occurs a ...
'' Emery, 1896 *''
Megalomyrmex mondabora ''Megalomyrmex mondabora'' is a Neotropical species of ants in the subfamily Myrmicinae. Known from Costa Rica. In Costa Rica this species occurs in wet forest habitats, typically in mature rainforest. Collections are from sea level to 800 m ...
'' Brandão, 1990 *'' Megalomyrmex mondaboroides'' Longino, 2010 *'' Megalomyrmex myops'' Santschi, 1925 *'' Megalomyrmex nocarina'' Longino, 2010 *'' Megalomyrmex osadrifti'' Boudinot, Sumnicht & Adams, 2013 *'' Megalomyrmex pacova'' Brandão, 1990 *'' Megalomyrmex piriana'' Brandão, 1990 *'' Megalomyrmex poatan'' Brandão, 1990 *'' Megalomyrmex pusillus'' Forel, 1912 *'' Megalomyrmex reina'' Longino, 2010 *'' Megalomyrmex silvestrii'' Wheeler, 1909 *''
Megalomyrmex staudingeri ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically ...
'' Emery, 1890 *''
Megalomyrmex symmetochus ''Megalomyrmex symmetochus'' is a species of ant in the subfamily Myrmicinae. It is native to Panama. ''M. symmetochus'' was discovered by William M. Wheeler in late July 1924 in the fungus gardens of the attine '' Sericomyrmex amabilis'' of B ...
'' Wheeler, 1925 *'' Megalomyrmex tasyba'' Brandão, 1990 *'' Megalomyrmex timbira'' Brandão, 1990 *''
Megalomyrmex wallacei ''Megalomyrmex wallacei'' is a Neotropical species of ants in the subfamily Myrmicinae. ''Megalomyrmex wallacei'' can be found in Brazil (Amazonas (Brazilian state), Amazonas, Rondônia, Pará, Tocantins), Guyana, Colombia, Costa Rica. This spec ...
'' Mann, 1916 *''
Megalomyrmex wettereri ''Megalomyrmex wettereri'' is a Neotropical species of ants in the subfamily Myrmicinae. ''Megalomyrmex wettereri'' is known from two lowland rainforest sites: Barro Colorado Island in Panama and La Selva Biological Station in Costa Rica. Nest ...
'' Brandão, 2003 *''
Megalomyrmex weyrauchi ''Megalomyrmex'' is a genus of ant in the subfamily Myrmicinae. The genus is known only from the Neotropics, where some of the species are specialized parasites or predators of Attini. Description It is difficult to characterize morphologically. ...
'' Kempf, 1970


References

*Bolton, B. (1994) Identification Guide to the Ant Genera of the World.
Harvard University Press Harvard University Press (HUP) is a publishing house established on January 13, 1913, as a division of Harvard University, and focused on academic publishing. It is a member of the Association of American University Presses. After the retirem ...
, Cambridge, MA, London, 222 pp. *Bolton, B. (2003) Synopsis and classification of Formicidae. ''Memoirs of the American Entomological Institute'' 71: 1–370 * * * * * *


External links

* {{Taxonbar, from=Q6808513 Myrmicinae Ant genera Taxonomy articles created by Polbot