Marajó Várzea
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The Marajó várzea (NT0138) is an
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...
of seasonally and tidally flooded
várzea forest A várzea forest is a seasonal floodplain forest inundated by whitewater rivers that occurs in the Amazon biome. Until the late 1970s, the definition was less clear and várzea was often used for all periodically flooded Amazonian forests. Althoug ...
in the
Amazon biome The Amazon biome ( pt, Bioma Amazônia) contains the Amazon rainforest, an area of tropical rainforest, and other ecoregions that cover most of the Amazon basin and some adjacent areas to the north and east. The biome contains blackwater and white ...
. It covers a region of sedimentary islands and floodplains at the mouth of the Amazon that is flooded twice daily as the ocean tides push the river waters onto the land. The flooded forests provide food for a wide variety of fruit-eating fish, aquatic mammals, birds and other fauna. It has no protected areas and is threatened by cattle and water-buffalo ranching, logging and fruit plantations.


Location

The Marajó várzea is at the mouth of the
Amazon River The Amazon River (, ; es, Río Amazonas, pt, Rio Amazonas) in South America is the largest river by discharge volume of water in the world, and the disputed longest river system in the world in comparison to the Nile. The headwaters of t ...
. It covers coastal areas of the states of
Pará Pará is a Federative units of Brazil, state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of Amapá, Maranhão, Tocantins (state), Tocantins, Mato Grosso, Amazonas (Brazilian state) ...
and
Amapá Amapá () is one of the 26 states of Brazil. It is in the northern region of Brazil. It is the second least populous state and the eighteenth largest by area. Located in the far northern part of the country, Amapá is bordered clockwise by Fr ...
, with an area of . Water levels are affected by freshwater flowing down the river and by tidal flows from the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
. The
várzea forest A várzea forest is a seasonal floodplain forest inundated by whitewater rivers that occurs in the Amazon biome. Until the late 1970s, the definition was less clear and várzea was often used for all periodically flooded Amazonian forests. Althoug ...
in the ecoregion starts where the
Xingu River The Xingu River ( ; pt, Rio Xingu, ; Mẽbêngôkre: ''Byti'', ) is a river in north Brazil. It is a southeast tributary of the Amazon River and one of the largest clearwater rivers in the Amazon basin, accounting for about 5% of its water. ...
joins the Amazon, which begins to broaden out. It covers the west of Marajó Island and many smaller islands in the Amazon channel, as well as parts of the mainland to the north and south of the river mouth including a small area of
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
. The nutrient-rich sediments from the river are deposited on the islands. The Marajó várzea ecoregion adjoins the
Xingu–Tocantins–Araguaia moist forests The Xingu–Tocantins–Araguaia moist forests (NT0180) is an ecoregion in the eastern Amazon basin. It is part of the Amazon biome. The ecoregion is one of the most severely degraded of the Amazon region, suffering from large-scale deforestation ...
and the
Tocantins–Araguaia–Maranhão moist forests The Tocantins–Araguaia–Maranhão moist forests (NT0170), also called the Tocantins/Pindaré moist forests, is an ecoregion in the north of Brazil to the south of the mouth of the Amazon River. It is part of the Amazon biome. The ecoregion con ...
to the south, and the
Uatuma–Trombetas moist forests The Uatuma–Trombetas moist forests (NT0173) is an ecoregion in northwest Brazil in the Amazon biome. It covers the Amazon basin north of the Amazon River from close to the Atlantic Ocean to the Rio Negro west of Manaus. The ecoregion is relativ ...
and an area of
Guianan savanna The Guianan savanna (NT0707) is an ecoregion in the south of Venezuela, Guyana and Suriname and the north of Brazil. It is in the Amazon biome. The savanna covers an area of rolling upland plains on the Guiana Shield between the Amazon and Orinoco ...
to the north. The
Gurupa várzea The Gurupa várzea (NT0126) is an ecoregion of seasonally and tidally flooded várzea forest along the Amazon River in the Amazon biome. Location The Gurupa várzea ecoregion is named after the Ilha Grande de Gurupá, an alluvial island in the mo ...
is upstream along the Amazon. There are strips of Amazon–Orinoco–Southern Caribbean mangroves along the coast. The
Amapá mangroves The Amapá mangroves (NT1402) is an ecoregion along the Atlantic coast of the state of Amapá in Brazil. The low coastal plain has been formed from recent sedimentation, including sediments deposited by the rivers and sediments carried northward f ...
are found to the north of the Amazon's mouth and the
Pará mangroves The Pará mangroves (NT1427) is an ecoregion along the Atlantic coast of the state of Pará in Brazil. They constitute the western extension of the Maranhão mangroves ecoregion. The mangroves are relatively intact, although they are under some pre ...
to the south.


Physical

The Marajó várzea covers flooded land at the mouth of the Amazon River. There are many islands including the
Marajó Marajó () is a large coastal island in the state of Pará, Brazil. It is the main and largest of the islands in the Marajó Archipelago. Marajó Island is separated from the mainland by Marajó Bay, Pará River, smaller rivers (especially ...
island and the smaller Porcos, Pará, Mututí and Uituquara islands. The estuary is a recent lowland that has been forming during the present
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
epoch, and is surrounded by
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
deposits. The constant action of tides and river flow form a maze of channels. Twice daily the tides push river discharge onto the low-lying land to a depth of . Soils in the many islands formed from the Amazon sediments are typically mottled clay with poor drainage. The ecoregion also includes slightly higher ground that is usually not flooded. The east of Marajó island is made of older Tertiary sediments. It is covered by flooded savanna and humid terra firme forest.


Climate

The
Köppen climate classification The Köppen climate classification is one of the most widely used climate classification systems. It was first published by German-Russian climatologist Wladimir Köppen (1846–1940) in 1884, with several later modifications by Köppen, notabl ...
is "Am": equatorial with monsoon rainfall. Average annual temperatures are about , ranging from about Average annual precipitation is about . Monthly rainfall varies from in October to over in March.


Ecology

The ecoregion is in the
Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
and the
tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discont ...
biome.


Flora

Vegetation is dominated by palms, and is shorter and less diverse than in surrounding areas. The most common palm species are murumuru (''
Astrocaryum murumuru ''Astrocaryum murumuru'' (Portuguese common name: murumuru) is a palm native to Amazon Rainforest vegetation in Brazil, which bears edible fruits. ''Murumuru'' butter, extracted from the seeds of the plant, may be used as a moisturizer.One re ...
''), raffia (''
Raphia taedigera ''Raphia taedigera'' is a palm species in the family Arecaceae, colloquially known as yolilla in Central American Spanish. It is found in parts of Western Africa, Central America and South America South America is a continent entirely ...
''), açaí (''Euterpe oleracea''), maripa (''
Attalea maripa ''Attalea maripa'', commonly called maripa palm is a palm native to tropical South America and Trinidad and Tobago. It grows up tall and can have leaves or fronds long. This plant has a yellow edible fruit which is oblong ovoid and cream. An e ...
''), bacaba (''
Oenocarpus bacaba ''Oenocarpus bacaba'' is an economically important monoecious fruiting palm native to South America and the Amazon Rainforest, which has edible fruits. This plant is cited in Flora Brasiliensis by Carl Friedrich Philipp von Martius. It can reach ...
''), patauá (''
Oenocarpus bataua ''Oenocarpus bataua'', the patawa, sehe, hungurahua (Ecuador) or mingucha, is a palm tree native to the Amazon rainforest. The tree produces edible fruits rich in high-quality oil.Vallejo Rendón, Darío 2002. "Oenocarpus bataua, seje"; ''Colombi ...
''), buriti (''
Mauritia flexuosa ''Mauritia flexuosa'', known as the moriche palm, ''ité'' palm, ''ita'', ''buriti'', ''muriti'', ''miriti'' (Brazil), ''canangucho'' (Colombia), ''acho'' (Ecuador), or ''aguaje'' (Peru), is a Arecaceae, palm tree. It grows in and near swamps and ...
''), ubuçu (''
Manicaria saccifera ''Manicaria'' is a palm genus which is found in Trinidad, Central and South America. It contains two recognized species: #''Manicaria martiana'' Burret – Colombia, northwestern Brazil #''Manicaria saccifera'' Gaertn. – Central America, Trin ...
''), cashapona (''
Socratea exorrhiza ''Socratea exorrhiza'', the walking palm or cashapona, is a Arecaceae, palm native to rainforests in tropical Central America, Central and South America. It can grow to 25 metres in height, with a stem diameter of up to 16 cm, but is more ty ...
'') and several species of Geonoma palms. There are large stands of the commercially valuable buriti and acai palms. Among other common trees in the tidal várzea are the timber species ''
Virola surinamensis ''Virola surinamensis'', known commonly as baboonwood, ucuuba, ucuhuba and ''chalviande'',Americas Regional Workshop (Conservation & Sustainable Management of Trees, Costa Rica, November 1996). 1998''Virola surinamensis''.The IUCN Red List of Thr ...
'', '' Cedrelinga castanaeformis'', ''
Ceiba pentandra ''Ceiba pentandra'' is a tropical tree of the order Malvales and the family Malvaceae (previously emplaced in the family Bombacaceae), native to Mexico, Central America and the Caribbean, northern South America, and (as the variety ''C. pentandr ...
'', '' Calycophyllum brasiliensis'' and other large trees such as
Ficus ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extending in ...
species, ''
Macrolobium acaciifolium ''Macrolobium'' is a legume genus in the subfamily Detarioideae. It is a tropical genus with about 80 species. Half occur in Brazil, where they are common in the floodplains of the Amazonian Basin. Members of the genus are used as ornamentals an ...
'', ''
Pachira aquatica ''Pachira aquatica'' is a tropical wetland tree in the mallow family Malvaceae, native to Central and South America where it grows in swamps. It is known by its common names Malabar chestnut, French peanut, Guiana chestnut, Provision tree, Saba n ...
'', ''
Symphonia globulifera ''Symphonia globulifera'', commonly known as boarwood, is a timber tree abundant in Central America, the Caribbean, South America and Africa. This plant is also used as a medicinal plant and ornamental plant. Common names Common trade names of t ...
'', '' Triplaris surinamensis'' and ''
Mora paraensis Mora may refer to: People * Mora (surname) Places Sweden * Mora, Säter, Sweden * Mora, Sweden, the seat of Mora Municipality * Mora Municipality, Sweden United States * Mora, Louisiana, an unincorporated community * Mora, Minnesota, a city * ...
''. The shrub '' Machaerium lanatum'' forms dense thickets along the banks of the rivers. There are large lianas in the flooded areas such as '' Strychnos blackii'', '' Landolphia paraensis'' and '' Guatteria scandens''. The ecoregion also includes seasonally flooded forest and permanently flooded
igapó Igapó (, from Old Tupi: "root forest") is a word used in Brazil for blackwater-flooded forests in the Amazon biome. These forests and similar swamp forests are seasonally inundated with freshwater. They typically occur along the lower reaches of ...
swamp forest.


Fauna

The várzea forest provides a mechanism for rapid capture and release of nutrients, and hosts many species of freshwater fish and aquatic mammals. There is greater diversity of fauna than in surrounding areas. 99 species of mammals have been reported. Big cats include
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
(''Panthera onca'') and
cougar The cougar (''Puma concolor'') is a large Felidae, cat native to the Americas. Its Species distribution, range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mamm ...
(''Puma concolor''). Fruiting trees such as the buriti palm are sources of food for grazing animals such as
gray brocket The gray brocket (''Mazama gouazoubira''), also known as the brown brocket, is a species of brocket deer from northern Argentina, Bolivia, southern Peru, eastern and southern Brazil, Paraguay, and Uruguay. It formerly included the Amazonian br ...
(''Mazama gouazoubira''),
red brocket The red brocket (''Mazama americana'') is a species of brocket deer from forests in South America, ranging from northern Argentina to Colombia and the Guianas. It also occurs on the Caribbean island of Trinidad (it also occurred on the island of ...
(''Mazama americana''), and
capybara The capybaraAlso called capivara (in Brazil), capiguara (in Bolivia), chigüire, chigüiro, or fercho (in Colombia and Venezuela), carpincho (in Argentina, Paraguay and Uruguay) and ronsoco (in Peru). or greater capybara (''Hydrochoerus hydro ...
(''Hydrochoerus hydrochaeris''). The estuary is home to large fish that swim through the flooded forest where they eat and spread fruit from the trees. Fish include pacus (
Metynnis ''Metynnis'' is a genus of serrasalmid fish from tropical and subtropical South America. They are herbivorous or omnivorous, and inhabit a wide range of freshwater habitats, ranging from rivers and streams (both fast- and slow-flowing), to flood ...
and
Mylossoma ''Mylossoma'' is a genus of serrasalmids from tropical and subtropical South America, including the basins of the Amazon, Orinoco, Lake Maracaibo and Paraguay- Paraná. These common fish are found both in main river sections and floodplains. T ...
genera),
tambaqui The tambaqui (''Colossoma macropomum'') is a large species of freshwater fish in the family Serrasalmidae. It is native to tropical South America, but kept in aquaculture and introduced elsewhere. It is also known by the names black pacu, blac ...
(''Colossoma macropomum''), pirarucus (''
Arapaima gigas ''Arapaima gigas'', also known as pirarucu, is a species of arapaima native to the basin of the Amazon River. Once believed to be the sole species in the genus, it is among the largest freshwater fish. The species is an obligate air-breather, so ...
''), and sardinhas (''
Triportheus angulatus ''Triportheus'' is a genus of characiform fishes from South America, including Trinidad, ranging from the Rio de la Plata basin to the basins of the Orinoco and Magdalena. Some are migratory. The largest species is up to in standard length ...
''). Endangered species include the
white-cheeked spider monkey The white-cheeked spider monkey (''Ateles marginatus'') is a species of spider monkey, a type of New World monkey, endemic to Brazil. It moves around the forest canopy in small family groups of two to four, part of larger groups of a few dozen an ...
(''Ateles marginatus''),
black bearded saki The black bearded saki (''Chiropotes satanas'') is a species of New World monkey, native to the Amazon rainforest of South America, specifically to an area of north-eastern Brazil. It is one of five species of bearded saki. Bearded sakis are med ...
(''Chiropotes satanas''),
giant otter The giant otter or giant river otter (''Pteronura brasiliensis'') is a South American carnivorous mammal. It is the longest member of the weasel family, Mustelidae, a globally successful group of predators, reaching up to . Atypical of musteli ...
(Pteronura brasiliensis),
green sea turtle The green sea turtle (''Chelonia mydas''), also known as the green turtle, black (sea) turtle or Pacific green turtle, is a species of large sea turtle of the family Cheloniidae. It is the only species in the genus ''Chelonia''. Its range exten ...
(''Chelonia mydas'') and
hawksbill sea turtle The hawksbill sea turtle (''Eretmochelys imbricata'') is a critically endangered sea turtle belonging to the family Cheloniidae. It is the only extant species in the genus ''Eretmochelys''. The species has a global distribution, that is largel ...
(''Eretmochelys imbricata''). The endemic
short-tailed opossum ''Monodelphis'' is a genus of marsupials in the family Didelphidae, commonly referred to as short-tailed opossums. They are found throughout South America. , the most recently described species is ''M. vossi''. Species * Sepia short-tailed o ...
(''Monodelphis maraxina'') is endangered. Other endemic mammals are the
seven-banded armadillo The seven-banded armadillo (''Dasypus septemcinctus''), also known as the Brazilian lesser long-nosed armadillo, is a species of armadillo from South America found in Paraguay, Argentina, Bolivia and Brazil. It is a solitary nocturnal, terrest ...
(''Dasypus septemcinctus''),
Recife broad-nosed bat The Recife broad-nosed bat (''Platyrrhinus recifinus''), is a species of bat from South America. It is named for the city of Recife in Brazil, where it was first recorded by Oldfield Thomas in 1901. Previously considered "Vulnerable", the Recif ...
(Platyrrhinus recifinus), Brazilian funnel-eared bat (''Natalus espiritosantensis''), Greenhall's dog-faced bat (Cynomops greenhalli),
silvery marmoset The silvery marmoset (''Mico argentatus'') is a New World monkey that lives in the eastern Amazon Rainforest in Brazil. The fur of the silvery marmoset is colored whitish silver-grey except for a dark tail. Remarkable are its naked, flesh-color ...
(''Mico argentatus''),
red-handed tamarin The golden-handed tamarin (''Saguinus midas''), also known as the red-handed tamarin or Midas tamarin, is a New World monkey belonging to the family Callitrichidae. Distribution and habitat This species is native to wooded areas north of the Ama ...
(''Saguinus midas''),
Azara's night monkey Azara's night monkey (''Aotus azarae''), also known as the southern night monkey, is a night monkey species from South America. It is found in Argentina, Bolivia, Brazil, Peru and Paraguay. The species is monogamous, with the males providing a ...
(''Aotus azarae'') and
crab-eating fox The crab-eating fox (''Cerdocyon thous''), also known as the forest fox, wood fox, bushdog (not to be confused with the bush dog) or maikong, is an extant species of medium-sized canid endemic to the central part of South America since at least ...
(''Cerdocyon thous''). The ecoregion is home to the endangered
West Indian manatee The West Indian manatee (''Trichechus manatus''), also known as the North American manatee, is a large, aquatic mammal native to warm coastal areas of the Caribbean, from the eastern US to northern Brazil. Living alone or in herds, it feeds on un ...
(''Trichechus manatus''). There are at least 540 species of birds. Aquatic bird species include heron and egret of the
Egretta ''Egretta'' is a genus of medium-sized herons, mostly breeding in warmer climates. Representatives of this genus are found in most of the world, and the little egret, as well as being widespread throughout much of the Old World, has now started ...
and Ardea genera,
whistling duck The whistling ducks or tree ducks are a subfamily, Dendrocygninae, of the duck, goose and swan family of birds, Anatidae. In other taxonomic schemes, they are considered a separate family, Dendrocygnidae. Some taxonomists list only one genu ...
(Dendrocygna species),
sharp-tailed ibis The sharp-tailed ibis (''Cercibis oxycerca'') is a species of ibis native to open wet savannas in parts of northern South America. Taxonomy and systematics The sharp-tailed ibis is monotypic, being the only representative of the genus ''Cercibis ...
(''Cercibis oxycerca''),
Theristicus ''Theristicus'' is a genus of birds in the family Threskiornithidae. They are found in open, grassy habitats in South America. All have a long, decurved dark Beak, bill, relatively short reddish legs that do not extend beyond the tail in flight ( ...
species and
roseate spoonbill The roseate spoonbill (''Platalea ajaja'') is a gregarious wading bird of the ibis and spoonbill family, Threskiornithidae. It is a resident breeder in both South and North America. Taxonomy The roseate spoonbill is sometimes placed in its own ...
(''Platalea ajaja''). Endemic birds include white-bellied seedeater (''Sporophila leucoptera''),
grassland yellow finch The grassland yellow finch (''Sicalis luteola'') is a small passerine bird. Despite its name, it is not a finch, but is a seedeater. These were formerly united with the buntings and American sparrows in the Emberizidae, but are now known to be ...
(''Sicalis luteola''), chalk-browed mockingbird (''Mimus saturninus''), tropical pewee (''Contopus cinereus''),
rufous-throated antbird The rufous-throated antbird (''Gymnopithys rufigula'') is a species of bird in the family Thamnophilidae. It is found in Brazil, French Guiana, Guyana, Suriname, and Venezuela. Its natural habitat is subtropical or tropical moist lowland forests ...
(''Gymnopithys rufigula''),
black-breasted puffbird The black-breasted puffbird (''Notharchus pectoralis'') is a species of bird in the family Bucconidae, the puffbirds, nunlets, and nunbirds. It is found in Colombia, Ecuador, and Panama. Taxonomy and systematics The black-breasted puffbird ...
(''Notharchus pectoralis'') and
plain-bellied emerald The plain-bellied emerald (''Chrysuronia leucogaster'') is a species of hummingbird in the "emeralds", tribe Trochilini of subfamily Trochilinae. It is found in Brazil, the Guianas, and Venezuela.HBW and BirdLife International (2021) Handbook o ...
(''Amazilia leucogaster''). Endangered birds include the scaled spinetail (''Cranioleuca muelleri'') and
yellow-bellied seedeater The yellow-bellied seedeater (''Sporophila nigricollis'') is a species of bird in the family Thraupidae, formerly placed with the American sparrows in the Emberizidae. Taxonomy and systematics Hooded seedeater The hooded seedeater was a proposed ...
(''Sporophila nigricollis'').


Conservation status

The ecoregion is in a region of the Amazon at the mouth of the great waterway that has long been heavily used by humans. It is vulnerable, and has no protected areas. Cattle and water buffalo ranches have degraded natural grasslands and forests and caused loss of food sources for capybaras, manatees and fruteating fish. Commercial logging on Marajó island has almost eliminated some valuable native tree species. Stands of Açaí palm planted for their commercially valuable fruit have replaced stands of native buriti palm. Global warming will force tropical species to migrate uphill to find areas with suitable temperature and rainfall. Low, flat ecoregions such as the Marajó várzea are extremely vulnerable.


Notes


Sources

* * * * * * * {{DEFAULTSORT:Marajó várzea Ecoregions of Brazil Amazon biome Neotropical tropical and subtropical moist broadleaf forests