''Pinus peuce'' (Macedonian pine or Balkan pine) (

Serbo-Croatian Serbo-Croatian () – also called Serbo-Croat (), Serbo-Croat-Bosnian (SCB), Bosnian-Croatian-Serbian (BCS), and Bosnian-Croatian-Montenegrin-Serbian (BCMS) – is a South Slavic language and the primary language of Serbia, Croatia, Bosnia and ...

/ Macedonian: молика, molika;

Bulgarian Bulgarian may refer to: * Something of, from, or related to the country of Bulgaria * Bulgarians, a South Slavic ethnic group * Bulgarian language, a Slavic language * Bulgarian alphabet * A citizen of Bulgaria, see Demographics of Bulgaria * Bul ...

: бяла мура, Byala mura) is a species of

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...

native to the mountains of Macedonia,

Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...

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Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = M ...

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, the extreme southwest of

Serbia Serbia (, ; Serbian language, Serbian: , , ), officially the Republic of Serbia (Serbian language, Serbian: , , ), is a landlocked country in Southeast Europe, Southeastern and Central Europe, situated at the crossroads of the Pannonian Bas ...

, and the extreme north of

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,Farjon, A. (2005). ''Pines. Drawings and descriptions of the genus Pinus'', ed.2. Brill, Leiden .Gymnosperm Databas
''Pinus peuce''
/ref> growing typically at (600-) 1,000-2,200 (-2,300) m altitude. It often reaches the alpine tree line in this area. The mature size is up to 35–40 m height, and 1.5 m trunk diameter. However, the height of the tree diminishes strongly near the upper forest limit and may even obtain shrub sizes. It is a member of the

white pine ''Pinus'', the pines, is a genus of approximately 111 extant tree and shrub species. The genus is currently split into two subgenera: subgenus ''Pinus'' (hard pines), and subgenus ''Strobus'' (soft pines). Each of the subgenera have been further ...

group, ''

Pinus A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...

'' subgenus ''

Strobus ''Pinus'', the pines, is a genus of approximately 111 extant tree and shrub species. The genus is currently split into two subgenera: subgenus ''Pinus'' (hard pines), and subgenus ''Strobus'' (soft pines). Each of the subgenera have been furth ...

'', and like all members of that group, the leaves ('needles') are in fascicles (bundles) of five, with a deciduous sheath. They are 6–11 cm long. Its pine cones are mostly 8–16 cm long, occasionally up to 20 cm long, green at first, becoming yellow-brown when mature, with broad, flat to downcurved scales. The 6–7 mm long

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

s have a 2 cm wing and can be wind-dispersed, but are also very often dispersed by

spotted nutcracker The spotted nutcracker, Eurasian nutcracker, or simply nutcracker (''Nucifraga caryocatactes'') is a passerine bird slightly larger than the Eurasian jay. It has a much larger bill and a slimmer looking head without any crest. The feathering over ...

Use

Macedonian pine is one of the most valuable

conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...

species in the

Balkan Peninsula The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...

. Its durable wood is highly valued in

construction Construction is a general term meaning the art and science to form objects, systems, or organizations,"Construction" def. 1.a. 1.b. and 1.c. ''Oxford English Dictionary'' Second Edition on CD-ROM (v. 4.0) Oxford University Press 2009 and com ...

, furniture production,

wood-carving Wood carving is a form of woodworking by means of a cutting tool (knife) in one hand or a chisel by two hands or with one hand on a chisel and one hand on a mallet, resulting in a wooden figure or figurine, or in the sculptural ornamentation ...

and

cooper Cooper, Cooper's, Coopers and similar may refer to: * Cooper (profession), a maker of wooden casks and other staved vessels Arts and entertainment * Cooper (producers), alias of Dutch producers Klubbheads * Cooper (video game character), in ...

age. The tree is also exceptionally good at adapting to severe mountain climate conditions, which makes it a valuable species for

afforestation Afforestation is the establishment of a forest or stand of trees (forestation) in an area where there was no previous tree cover. Many government and non-governmental organizations directly engage in afforestation programs to create forests a ...

on high terrain for protection against

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is distin ...

. The local population use ''P. peuce''

resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...

to cure wounds,

pectoral Pectoral may refer to: * The chest region and anything relating to it. * Pectoral cross, a cross worn on the chest * a decorative, usually jeweled version of a gorget * Pectoral (Ancient Egypt), a type of jewelry worn in ancient Egypt * Pectoralis ...

, skin and stomach diseases,

varicose veins Varicose veins, also known as varicoses, are a medical condition in which superficial veins become enlarged and twisted. These veins typically develop in the legs, just under the skin. Varicose veins usually cause few symptoms. However, some indi ...

and other illnesses. Macedonian pine is also a popular ornamental tree in

park A park is an area of natural, semi-natural or planted space set aside for human enjoyment and recreation or for the protection of wildlife or natural habitats. Urban parks are urban green space, green spaces set aside for recreation inside t ...

s and large

garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate both ...

s, giving reliable steady though not fast growth on a wide range of sites. It is very tolerant of severe winter cold, hardy down to at least -45 °C, and also of wind exposure. It is locally

naturalised Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the i ...

Punkaharju Punkaharju is a former municipality of Finland. It was consolidated with the town of Savonlinna on January 1, 2013. It was located in the provinces of Finland, province of Eastern Finland and is part of the Southern Savonia regions of Finland, reg ...

in eastern Finland. Like other European and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

n white pines, Macedonian pine is very resistant to

white pine blister rust ''Cronartium ribicola'' is a species of rust fungus in the family Cronartiaceae that causes the disease white pine blister rust. Other names include: (French), (German), (Spanish). ''Cronartium ribicola'' is native to China, and was subsequ ...

(''Cronartium ribicola'').Søegaard, B. (1972). Relative blister rust resistance of native and introduced white pines in Europe. Pages 233–239 in Bingham, R. T. et al., eds. Biology of Rust Resistance in Forest Trees. ''U.S. Dept. of Agriculture Forest Service Misc. Publ''. 1221.Popnikola, N., M. Jovancevic, and M. Vidakovic. 1978. Genetics of Pinus peuce Gris. ''Annales Forestales'' 7/6: 187-206. This fungal disease was accidentally introduced from

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

into

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

, where it has caused severe mortality in the American native white pines (e.g.

western white pine Western white pine (''Pinus monticola''), also called silver pine and California mountain pine, is a species of pine in the family Pinaceae. It occurs in mountain ranges of northwestern North America. It is the state tree of Idaho. Description ...

sugar pine ''Pinus lambertiana'' (commonly known as the sugar pine or sugar cone pine) is the tallest and most massive pine tree, and has the longest cones of any conifer. The species name ''lambertiana'' was given by the Scottish botanist David Douglas, ...

whitebark pine ''Pinus albicaulis'', known by the common names whitebark pine, white bark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine ...

) in many areas. Macedonian pine is of great value for research into hybridisation and genetic modification to develop rust resistance in these species; hybrids with

eastern white pine ''Pinus strobus'', commonly called the eastern white pine, northern white pine, white pine, Weymouth pine (British), and soft pine is a large pine native to eastern North America. It occurs from Newfoundland, Canada west through the Great Lakes ...

inherit some resistance.

Other

Synonyms include ''Pinus cembra'' var. ''fruticosa'' Griseb., ''Pinus excelsa'' var. ''peuce'' (Griseb.) Beissn., ''Pinus peuce'' var. ''vermiculata'' Christ, and Balkan pine.Dirr, M. (1997). ''Dirr's Hardy trees and shrubs: an illustrated encyclopedia''. Timber Press , page. 282 File:Pinus peuce Habitus BotGardBln0906.jpg, Cultivated specimen, Berlin Botanical Gardens File:Pinus peuce cones.jpg, Foliage and cones Pinus peuce young female cone - Keila.jpg, Young female cone

Notes

References

External links

''Pinus peuce''
- information, genetic conservation units and related resources.

European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that supports the conservation and sustainable use of forest genetic resources in Europe. The programme’s tasks include to coordinate and promote ''in situ ' ...

(EUFORGEN) {{Authority control peuce Flora of Southeastern Europe