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''Myrica cerifera'' is a small
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
or large
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
native to North and Central America and the Caribbean. Its common names include southern wax myrtle, southern bayberry, candleberry, bayberry tree, and tallow shrub. It sees uses both in the garden and for
candlemaking A candle is an ignitable wick embedded in wax, or another flammable solid substance such as tallow, that provides light, and in some cases, a fragrance. A candle can also provide heat or a method of keeping time. A person who makes candles i ...
, as well as a
medicinal plant Medicinal plants, also called medicinal herbs, have been discovered and used in traditional medicine practices since prehistoric times. Plants synthesize hundreds of chemical compounds for various functions, including Plant defense against her ...
.


Description

''Myrica cerifera'' is a small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
or large
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
, reaching up to 14m tall. It is adaptable to many habitats, growing naturally in wetlands, near
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s and
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...
s,
sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, fl ...
s, fields, hillsides,
pine barren Pine barrens, pine plains, sand plains, or pineland areas occur throughout the U.S. from Florida to Maine (see Atlantic coastal pine barrens) as well as the Midwest, West, and Canada and parts of Eurasia. Perhaps the most well known pine-barr ...
s, and in both
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant ...
and mixed-broadleaf forests. ''M. cerifera'' can weather coastal storms, long droughts, and tropical high temperatures. In nature, it ranges from
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, northward into the southeastern and south-central United States. Wax Myrtle can be successfully cultivated as far north as the New York City area and southern Ohio Valley. It also grows in
Bermuda ) , anthem = "God Save the King" , song_type = National song , song = " Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , e ...
and the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
. In terms of succession, ''M. cerifera'' is often one of the first plants to colonize an area. ''M. cerifera'' is an evergreen. The
glandular In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are long, have a leathery texture and serrated edges, and contain aromatic compounds. The plant is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
, with male and female flowers borne in catkins on separate plants. Male flowers have three or four
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s, and are surrounded by short
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s. The female flowers develop into fruit, which are globular and surrounded by a natural
wax Waxes are a diverse class of organic compounds that are lipophilic, malleable solids near ambient temperatures. They include higher alkanes and lipids, typically with melting points above about 40 Â°C (104 Â°F), melting to giv ...
-like coating. The species flowers from late winter to spring, and bear fruit in late summer or fall. No
endosperm The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the embryo and ...
is present on the seeds. ''M. cerifera'' can also reproduce clonally through runners. This species occurs in two forms, but there is no clear dividing line between them, many intermediate forms occurring. Specimens in drier and sandier areas are shrub-like, have
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s and smaller leaves. Those growing in damper situations with richer soil are more tree-like with bigger leaves. The fruit is a source of food for many bird species, including the northern
bobwhite quail The northern bobwhite (''Colinus virginianus''), also known as the Virginia quail or (in its home range) bobwhite quail, is a ground-dwelling bird native to Canada, the United States, Mexico, and Cuba, with introduced populations elsewhere in th ...
and the
wild turkey The wild turkey (''Meleagris gallopavo'') is an Upland game bird, upland ground bird native to North America, one of two extant species of Turkey (bird), turkey and the heaviest member of the order Galliformes. It is the ancestor to the domestic ...
. In winter, the seeds are important foods for the
Carolina wren The Carolina wren (''Thryothorus ludovicianus'') is a common species of wren that is a resident in the eastern half of the United States of America, the extreme south of Ontario, Canada, and the extreme northeast of Mexico. Severe winters restri ...
and species of
tree sparrow The Eurasian tree sparrow (''Passer montanus'') is a passerine bird in the sparrow family with a rich chestnut crown and nape, and a black patch on each pure white cheek. The sexes are similarly plumaged, and young birds are a duller version o ...
. To a point, ''M. cerifera'' will also provide habitat for the northern bobwhite quail. Birds' digestive systems remove the wax from the fruit, a prerequisite for
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
. This plant's
root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
s possess
root nodule Root nodules are found on the roots of plants, primarily legumes, that form a symbiosis with nitrogen-fixing bacteria. Under nitrogen-limiting conditions, capable plants form a symbiotic relationship with a host-specific strain of bacteria known a ...
s, which harbor a symbiotic species of actinomycotal bacteria, which fixes nitrogen at a faster rate than do the
legumes A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock fo ...
. The above-ground growth of ''M. cerifera'' is often killed by
wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire ...
s unless the fire is very small or transient. In the latter situation, only the most recent primary growth may be incinerated. Because the leaves, stem, and branches contain flammable aromatic compounds, a specimen of ''M. cerifera'' is in fact a
fire hazard Fire safety is the set of practices intended to reduce the destruction caused by fire. Fire safety measures include those that are intended to prevent the ignition of an uncontrolled fire and those that are used to limit the development and ef ...
. In contrast to the flammability of its top growth, ''M. ceriferas
root system In mathematics, a root system is a configuration of vectors in a Euclidean space satisfying certain geometrical properties. The concept is fundamental in the theory of Lie groups and Lie algebras, especially the classification and representati ...
is fire-resistant. By 1991, no known fire had killed this plant's roots. However, this plant will not survive repeated destruction of its top growth indefinitely. Three consecutive years of burning may kill all plants affected. After less damaging fires, new shoots will regrow from below ground. This regrowth is most rapid in the first season after a fire.


Taxonomy

This plant is one of several ''
Myrica ''Myrica'' is a genus of about 35–50 species of small trees and shrubs in the family Myricaceae, order Fagales. The genus has a wide distribution, including Africa, Asia, Europe, North America and South America, and missing only from Austra ...
'' species that are sometimes
split Split(s) or The Split may refer to: Places * Split, Croatia, the largest coastal city in Croatia * Split Island, Canada, an island in the Hudson Bay * Split Island, Falkland Islands * Split Island, Fiji, better known as Hạfliua Arts, enterta ...
into the genus ''Morella'', e.g. in the
Integrated Taxonomic Information System The Integrated Taxonomic Information System (ITIS) is an American partnership of federal agencies designed to provide consistent and reliable information on the taxonomy of biological species. ITIS was originally formed in 1996 as an interagenc ...
. This species also has several
synonyms A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
aside from the ''Myrica''/''Morella'' split: ''Cerothamnus pumilus'', ''C. ceriferus'', ''Myrica cerifera var. pumila'', and ''Myrica pusilla''. ''Myrica cerifera'' is similar to '' M. pensylvanica'' and '' M. caroliniensis''. These plants' scent or
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s can distinguish them. The generic name ''
Myrica ''Myrica'' is a genus of about 35–50 species of small trees and shrubs in the family Myricaceae, order Fagales. The genus has a wide distribution, including Africa, Asia, Europe, North America and South America, and missing only from Austra ...
'' comes from a
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word ''myrike'', which refers to some fragrant plant (possibly
tamarisk The genus ''Tamarix'' (tamarisk, salt cedar, taray) is composed of about 50–60 species of flowering plants in the family Tamaricaceae, native to drier areas of Eurasia and Africa. The generic name originated in Latin and may refer to the Tam ...
). The specific name means "wax-bearing".


Uses


Food

The leaves can be washed, dried, and used to season stews or cooking sauces. The berries can also be used as seasoning.


Ornamental

''Myrica cerifera'' finds use in
gardening Gardening is the practice of growing and cultivating plants as part of horticulture. In gardens, ornamental plants are often grown for their flowers, foliage, or overall appearance; useful plants, such as root vegetables, leaf vegetables, fruits ...
and
horticulture Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
. It has been commonly grown in American
hardiness zone A hardiness zone is a geographic area defined as having a certain average annual minimum temperature, a factor relevant to the survival of many plants. In some systems other statistics are included in the calculations. The original and most wide ...
s of 11 to 7. '' M. pensylvanica'' substitutes for ''M. cerifera'' in areas colder than zone 6. Since the species is adaptable, it will tolerate many conditions, although it has a need for frequent
pruning Pruning is a horticultural, arboricultural, and silvicultural practice involving the selective removal of certain parts of a plant, such as branches, buds, or roots. The practice entails the ''targeted'' removal of diseased, damaged, dead, ...
. It can handle abuse from bad pruning, however. The species has at least four
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
s. Those dubbed Fairfax, Jamaica Road, and Don's Dwarf differ from the "typical" specimen in habit and form. The latter two are also resistant to
leaf spot A leaf spot is a limited, discoloured, diseased area of a leaf that is caused by fungal, bacterial or viral plant diseases, or by injuries from nematodes, insects, environmental factors, toxicity or herbicides. These discoloured spots or lesions ...
. Var. ''pumila'' is a dwarf cultivar.


Herbalism

Bayberry root bark has a history of use in
herbalism Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern reme ...
. The plant contains several organic compounds, including:
triterpene Triterpenes are a class of chemical compounds composed of three terpene units with the molecular formula C30H48; they may also be thought of as consisting of six isoprene units. Animals, plants and fungi all produce triterpenes, including squa ...
s such as myricadiol, taraxerol, and taraxerone, as well as chemicals such as different
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s,
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'', ...
s,
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...
s, gums, and
phenol Phenol (also called carbolic acid) is an aromatic organic compound with the molecular formula . It is a white crystalline solid that is volatile. The molecule consists of a phenyl group () bonded to a hydroxy group (). Mildly acidic, it req ...
s. Myricadiol has a slight impact on levels of
potassium Potassium is the chemical element with the symbol K (from Neo-Latin ''kalium'') and atomic number19. Potassium is a silvery-white metal that is soft enough to be cut with a knife with little force. Potassium metal reacts rapidly with atmosphe ...
and
sodium Sodium is a chemical element with the symbol Na (from Latin ''natrium'') and atomic number 11. It is a soft, silvery-white, highly reactive metal. Sodium is an alkali metal, being in group 1 of the periodic table. Its only stable iso ...
, while a substance called
myricitrin Myricitrin is a plant compound, the 3-O-α-L-rhamnopyranoside of myricetin. Occurrences It can be isolated from the root bark of ''Myrica cerifera'' (Bayberry, a small tree native to North America), in '' Myrica esculenta'', in '' Nymphaea lotus ...
has antibiotic properties. The
Choctaw The Choctaw (in the Choctaw language, Chahta) are a Native American people originally based in the Southeastern Woodlands, in what is now Alabama and Mississippi. Their Choctaw language is a Western Muskogean language. Today, Choctaw people are ...
boiled bayberry and used the result as a treatment for
fever Fever, also referred to as pyrexia, is defined as having a body temperature, temperature above the human body temperature, normal range due to an increase in the body's temperature Human body temperature#Fever, set point. There is not a single ...
s. In 1722, it was reported that colonists in Louisiana drank a mixture of wax and hot water to treat severe
dysentery Dysentery (UK pronunciation: , US: ), historically known as the bloody flux, is a type of gastroenteritis that results in bloody diarrhea. Other symptoms may include fever, abdominal pain, and a feeling of incomplete defecation. Complications ...
. Bayberry was reported in an account from 1737 as being used to treat
convulsions A convulsion is a medical condition where the body muscles contract and relax rapidly and repeatedly, resulting in uncontrolled shaking. Because epileptic seizures typically include convulsions, the term ''convulsion'' is sometimes used as a s ...
,
colic Colic or cholic () is a form of pain that starts and stops abruptly. It occurs due to muscular contractions of a hollow tube ( small and large intestine, gall bladder, ureter, etc.) in an attempt to relieve an obstruction by forcing content out ...
,
palsy Palsy is a medical term which refers to various types of paralysisDan Agin, ''More Than Genes: What Science Can Tell Us About Toxic Chemicals, Development, and the Risk to Our Children;; (2009), p. 172. or paresis, often accompanied by weakness a ...
, and
seizure An epileptic seizure, informally known as a seizure, is a period of symptoms due to abnormally excessive or synchronous neuronal activity in the brain. Outward effects vary from uncontrolled shaking movements involving much of the body with los ...
s. Starting in the early 19th century, the herbalist
Samuel Thomson Samuel Thomson (9 February 1769 – 5 October 1843) was a self-taught American herbalist and botanist, best known as the founder of the alternative system of medicine known as "Thomsonian Medicine", which enjoyed wide popularity in the United ...
recommended this plant for producing "heat" within the body and as a treatment for infectious diseases and
diarrhea Diarrhea, also spelled diarrhoea, is the condition of having at least three loose, liquid, or watery bowel movements each day. It often lasts for a few days and can result in dehydration due to fluid loss. Signs of dehydration often begin wi ...
. That use of bayberry waned later in the 19th century, in favor of using it for a variety of ailments, including a
topical A topical medication is a medication that is applied to a particular place on or in the body. Most often topical medication means application to body surfaces such as the skin or mucous membranes to treat ailments via a large range of classes ...
use for bleeding
gums The gums or gingiva (plural: ''gingivae'') consist of the mucosal tissue that lies over the mandible and maxilla inside the mouth. Gum health and disease can have an effect on general health. Structure The gums are part of the soft tissue lin ...
. For twenty years starting in 1916, bayberry root bark was listed in the American
National Formulary A formulary is a list of pharmaceutical drugs, often decided upon by a group of people, for various reasons such as insurance coverage or use at a medical facility. Traditionally, a formulary contained a collection of formulas for the compoundin ...
. Use of bayberry in herbalism has declined since its peak in popularity in the 19th century. The plant is still used today in the treatment of fever, diarrhea, and a few other ailments. The chemical myricitrin has anti-fever properties. In addition, that chemical, along with the tannins, has anti-diarrheal properties. Myricitrin works as an antibiotic, while the tannins have
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin ''adstringere'', which means "to bind fast". Calamine lotion, witch hazel, and yerba mansa, a Californian plant ...
properties. In general, either a
decoction Decoction is a method of extraction by boiling herbal or plant material (which may include stems, roots, bark and rhizomes) to dissolve the chemicals of the material. It is the most common preparation method in various herbal-medicine systems. Dec ...
or a
tincture A tincture is typically an extract of plant or animal material dissolved in ethanol (ethyl alcohol). Solvent concentrations of 25–60% are common, but may run as high as 90%.Groot Handboek Geneeskrachtige Planten by Geert Verhelst In chemistr ...
is used.
Infusion Infusion is the process of extracting chemical compounds or flavors from plant material in a solvent such as water, oil or alcohol, by allowing the material to remain suspended in the solvent over time (a process often called steeping). An inf ...
s and a topical paste have also been used.
Pregnant Pregnancy is the time during which one or more offspring develops (gestation, gestates) inside a woman, woman's uterus (womb). A multiple birth, multiple pregnancy involves more than one offspring, such as with twins. Pregnancy usually occur ...
women should not use bayberry. In addition,
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'', ...
action relating to cancer is unclear, with studies indicating both pro and anti-cancer effects. Bayberry, just like any other medicinal plant, should only be used under the supervision of a physician.


Candles

Southern bayberry's fruits are a traditional source of the
wax Waxes are a diverse class of organic compounds that are lipophilic, malleable solids near ambient temperatures. They include higher alkanes and lipids, typically with melting points above about 40 Â°C (104 Â°F), melting to giv ...
for old-fashioned
Christmas Christmas is an annual festival commemorating Nativity of Jesus, the birth of Jesus, Jesus Christ, observed primarily on December 25 as a religious and cultural celebration among billions of people Observance of Christmas by country, around t ...
decorations called bayberry
candle A candle is an ignitable wick embedded in wax, or another flammable solid substance such as tallow, that provides light, and in some cases, a fragrance. A candle can also provide heat or a method of keeping time. A person who makes candles i ...
s. The wax was extracted by boiling the berries, and skimming off the floating hydrocarbons. The fats were then boiled again and then strained. After that the liquid was usable in
candle making Candle making was developed independently in many places throughout history. Dipped candles made from tallow were made by the Romans beginning about 1000 BCE. Evidence for candles made from whale fat in China dates back to the Qin Dynasty (221â ...
, whether through dipping or molding. Southern bayberry is not the only plant usable for making bayberry candles, however. Its close relatives are also usable. Southern bayberry and its relatives have largely been supplanted in candlemaking by substitutes made from
paraffin Paraffin may refer to: Substances * Paraffin wax, a white or colorless soft solid that is used as a lubricant and for other applications * Liquid paraffin (drug), a very highly refined mineral oil used in cosmetics and for medical purposes * Alkan ...
. The substitute candles have
artificial color Food coloring, or color additive, is any dye, pigment, or substance that imparts color when it is added to food or drink. They come in many forms consisting of liquids, powders, gels, and pastes. Food coloring is used in both commercial food ...
s and scents that create candles that look and smell similar to natural ones.


Soap

Wax extracted from boiling the fruit of ''Myrica cerifera'' was also used to make scented bayberry soap by early North-American settlers.


References

{{Taxonbar, from=Q1109693 cerifera Plants used in traditional Native American medicine Trees of the United States Flora of North America Flora of Central America Plants described in 1753 Taxa named by Carl Linnaeus Flora of the Caribbean Taxobox binomials not recognized by IUCN