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''Myosotis lyallii'' subsp. ''elderi'' is a subspecies of flowering plant in the family Boraginaceae, endemic to New Zealand. Lucy Moore described ''Myosotis elderi'' in 1961, and Heidi Meudt and Jessie Prebble treated it as a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of ''M. lyallii'' in 2018. Plants of this subspecies of forget-me-not are perennial with a prostrate habit, bracteate inflorescences, and white corollas with partially exserted anthers.


Taxonomy and etymology

''Myosotis lyallii'' subsp. ''elderi'' (
L.B.Moore Lucy Beatrice Moore (14 July 1906 – 9 June 1987) was a New Zealand botanist and ecologist. Biography Early life and education Moore was born in Warkworth, New Zealand, on 14 July 1906, the daughter of Janet Morison and Harry Blomfield Moor ...
) Meudt & Prebble is in the plant family Boraginaceae. The species ''Myosotis elderi'' was described in 1961 by Lucy Moore. In 2018, it was treated as a subspecies of ''M. lyallii.'' The holotype specimen of ''Myosotis elderi'' is lodged at herbarium WELT at the
Museum of New Zealand Te Papa Tongarewa The Museum of New Zealand Te Papa Tongarewa is New Zealand's national museum and is located in Wellington. ''Te Papa Tongarewa'' translates literally to "container of treasures" or in full "container of treasured things and people that spring fr ...
(WELT SP002648). The specific epithet, ''elderi'', was chosen to honor
Norman Elder Norman Sam Elder (July 17, 1939 – October 15, 2003) was a Canadian explorer, exotic animal owner, writer, artist, Olympic equestrian and one of Toronto's eccentrics. He was the owner of the Norman Elder Museum at 140 Bedford Road in th ...
(1896–1974), a New Zealand teacher, engineer and botanist. This is one of two subspecies recognized in '' M. lyallii;'' the other is ''M. lyallii'' subsp. ''lyallii''''.'' The subspecies are largely
allopatric Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
, as ''M. lyallii'' subsp. ''elderi'' is generally found on the dry, eastern side of the main divide of the South Island, whereas ''M. lyallii'' subsp. ''lyallii'' is generally found in the wet, western side.'''' In ''M. lyallii'' subsp. ''elderi,'' the anthers are at least partly below the faucal scales (partially exserted), whereas in ''M. lyallii'' subsp. ''lyallii,'' they are usually wholly above the faucal scales (fully exserted).'''' Furthermore, ''M. lyallii'' subsp. ''elderi'' has shorter filaments (< 0.6 mm) and wholly bracteate inflorescences, whereas ''M. lyallii'' subsp. ''lyallii'' has longer filaments > 0.6 mm long and at least some inflorescences that are partially bracteate.''''


Phylogeny

To date, ''Myosotis lyallii'' subsp. ''elderi'' has not been included in any published phylogenetic analyses using standard DNA sequencing markers (
nuclear ribosomal DNA Nuclear may refer to: Physics Relating to the Atomic nucleus, nucleus of the atom: *Nuclear engineering *Nuclear physics *Nuclear power *Nuclear reactor *Nuclear weapon *Nuclear medicine *Radiation therapy *Nuclear warfare Mathematics *Nuclear ...
and chloroplast DNA regions).


Description

''Myosotis lyallii'' subsp. ''elderi'' plants are rosettes that grow together forming loose clumps. The rosette leaves have petioles usually 1–6 mm long (rarely up to 13 mm long). The rosette leaf blades are 3–21 mm long by 2–13 mm wide (length: width ratio 1.1–3.6: 1), usually narrowly obovate to broadly obovate, usually widest at or above the middle, and usually with an obtuse apex. The upper surface of the leaf are densely covered in long, flexuous (sometimes curved), usually patent, antrorse (forward-facing) hairs that are oriented parallel to the mid vein; the leaf margin and petiole margin have patent to erect hairs. The lower surface of the leaf is similar except it can be glabrous or with isolated hairs on the midrib only. Each rosette has 1–19 prostrate or ascending, unbranched or once-branched, wholly bracteate inflorescences that are up to 105 mm long (rarely up to 230 mm long). The cauline leaves (sometimes called bracts) are similar to the rosette leaves but smaller, and decrease in size and become sessile toward the tip. Each inflorescence has 2–15 flowers, each borne on a short
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
, and each usually with a bract. The calyx is 3–7 mm long at flowering and 4–8 mm long at fruiting, lobed to one-third to two-thirds its length, and densely covered in long, antrorse, mostly flexuous, appressed to patent hairs (sometimes mixed with retrorse or backward-facing hairs, or curved or rarely hooked hairs, on the calyx base). The corolla is white, up to 11 mm in diameter, with a cylindrical tube, petals that are broadly to very broadly ovate or obovate to very broadly obovate, and flat, and small yellow scales alternating with the petals. The anthers are 2–9 mm long and partially exserted, with the tips or upper third only surpassing the scales. The filaments are 0–0.5 mm long. The four smooth, shiny, light, medium or dark brown nutlets are 1.1–2.4 mm long by 0.6–1.5 mm wide and narrowly ovoid to ovoid in shape. The chromosome number of ''M. lyallii'' subsp. ''elderi'' is unknown. ''M. lyallii'' subsp. ''elderi'' has ''M. australis'' type pollen. It flowers and fruits from November–April, with the main flowering period December–February and the main fruiting period January–March.


Distribution and habitat

''Myosotis lyallii'' subsp. ''elderi'' is a forget-me-not endemic to the mountains of eastern areas of the South Island New Zealand from 1100–2200 m ASL in the following ecological districts: Marlborough, Westland, Canterbury, Otago, Southland and Fiordland. It is also found on the North Island in the Southern North Island ecological district. ''M. lyallii'' subsp. ''elderi'' is an occasional plant in rocky, exposed, are rather bare high-elevation sites such as saddles, ridges and moraines, in sparsely vegetated fellfields and
tussock Tussock may refer to: * Tussock grass, a group of species in the family Poaceae *Floating island * Lymantriinae, called tussock moths or tussocks See also * Hassock (disambiguation) Hassock may refer to: * Kneeler, a cushion or a piece of f ...
-grasslands, herbfields and cushion turf.


Conservation status

''Myosotis lyallii'' subsp. ''elderi'' is listed (as ''M. elderi'') as "Threatened - Nationally Vulnerable", with the qualifiers "DP" (Data Poor) and "Sp" (Sparse) in the most recent assessment (2017-2018) under the New Zealand Threatened Classification system for plants.


References


External links


''Myosotis lyallii subsp. elderi'' occurrence data from Australasian Virtual Herbarium
{{Taxonbar, from=Q89179621 lyallii subsp. elderi Endemic flora of New Zealand Endangered flora of New Zealand Plants described in 1961