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The muskellunge ''(Esox masquinongy)'', often shortened to muskie, musky or lunge is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of large
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
predatory fish Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, pike/muskellunge, walleye, perch and salmon. Some omnivorous fish, such as the red-bellied piranha ...
native to
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. It is the largest member of the
pike Pike, Pikes or The Pike may refer to: Fish * Blue pike or blue walleye, an extinct color morph of the yellow walleye ''Sander vitreus'' * Ctenoluciidae, the "pike characins", some species of which are commonly known as pikes * ''Esox'', genus of ...
family,
Esocidae Esocidae is a family of fish in the order Esociformes, which contains pike, pickerel, and mudminnows. While the family traditionally only contained the genus ''Esox'', recent genetic and paleontological research have recovered ''Novumbra'' and ' ...
.


Origin of name

The name "muskellunge" originates from the
Ojibwe The Ojibwe, Ojibwa, Chippewa, or Saulteaux are an Anishinaabe people in what is currently southern Canada, the northern Midwestern United States, and Northern Plains. According to the U.S. census, in the United States Ojibwe people are one of ...
words ''maashkinoozhe'' (meaning "great fish"), ''maskinoše'' or ''mashkinonge'' (meaning "big pike" or "ugly pike") and the
Algonquin Algonquin or Algonquian—and the variation Algonki(a)n—may refer to: Languages and peoples *Algonquian languages, a large subfamily of Native American languages in a wide swath of eastern North America from Canada to Virginia **Algonquin la ...
word ''maskinunga'', which are borrowed into the
Canadian French Canadian French (french: français canadien) is the French language as it is spoken in Canada. It includes Varieties of French#Canada, multiple varieties, the most prominent of which is Quebec French, Québécois (Quebec French). Formerly ''Can ...
words ''masquinongé'' or ''maskinongé''. In
English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ide ...
, before settling on the common name "muskellunge", there have been at least 94 common names applied to this species, including but not limited to: ''muskelunge'', ''muscallonge'', ''muskallonge'', ''milliganong'', ''maskinonge'', ''maskalonge'', ''mascalonge'', ''maskalung'', ''muskinunge'' and ''masquenongez''. The word muskellunge is German and means "muscle lung".


Description

Muskellunge closely resemble other
esocid ''Esox'' is a genus of freshwater fish commonly known as pike or pickerel. It is the type genus of the family Esocidae. The type species of the genus is ''Esox lucius'', the northern pike. ''Esox'' has been present in Laurentia (which late ...
s such as the
northern pike The northern pike (''Esox lucius'') is a species of carnivorous fish of the genus '' Esox'' (the pikes). They are typical of brackish and fresh waters of the Northern Hemisphere (''i.e.'' holarctic in distribution). They are known simply as a ...
and
American pickerel The American pickerels are two subspecies of ''Esox americanus'', a medium-sized species of North American freshwater predatory fish belonging to the pike family (genus ''Esox'' in family Esocidae of order Esociformes): * Redfin pickerel, somet ...
in both appearance and behavior. Like the northern pike and other aggressive pikes, the body plan is typical of
ambush predator Ambush predators or sit-and-wait predators are carnivorous animals that capture or trap prey via stealth, luring or by (typically instinctive) strategies utilizing an element of surprise. Unlike pursuit predators, who chase to capture prey us ...
s with an elongated body, flat head, and dorsal, pelvic, and anal
fins A fin is a thin component or appendage attached to a larger body or structure. Fins typically function as foils that produce lift or thrust, or provide the ability to steer or stabilize motion while traveling in water, air, or other fluids. Fin ...
set far back on the body. Muskellunge are typically long and weigh , though some have reached up to and almost . Martin Arthur Williamson caught a muskellunge with a weight of 61.25 lb (27.8 kg) in November 2000 on
Georgian Bay Georgian Bay (french: Baie Georgienne) is a large bay of Lake Huron, in the Laurentia bioregion. It is located entirely within the borders of Ontario, Canada. The main body of the bay lies east of the Bruce Peninsula and Manitoulin Island. To ...
. The fish are a light silver, brown, or green, with dark vertical stripes on the flank, which may tend to break up into spots. In some cases, markings may be absent altogether, especially in fish from
turbid Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of water quality. Fluids can ...
waters. This is in contrast to northern pike, which have dark bodies with light markings. A reliable method to distinguish the two similar species is by counting the sensory pores on the underside of the
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
. A muskie will have seven or more per side, while the northern pike never has more than six. The lobes of the caudal (tail) fin in muskellunge come to a sharper point, while those of northern pike are more generally rounded. In addition, unlike pike, muskies have no scales on the lower half of their opercula. Anglers seek large muskies as trophies or for sport. In places where muskie are not native, such as in Maine, anglers are encouraged not to release the fish back into the water because of their negative impact on native populations of trout and other smaller fish species.


Habitat

Muskellunge are found in oligotrophic and
mesotrophic lake The Trophic State Index (TSI) is a classification system designed to rate water bodies based on the amount of biological productivity they sustain. Although the term "trophic index" is commonly applied to lakes, any surface water body may be inde ...
s and large
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s from northern
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and the ...
, northern
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...
, and northern
Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over to ...
through the
Great Lakes region The Great Lakes region of North America is a binational Canadian–American region that includes portions of the eight U.S. states of Illinois, Indiana, Michigan, Minnesota, New York, Ohio, Pennsylvania and Wisconsin along with the Canadian p ...
,
Chautauqua Lake Chautauqua Lake is located entirely within Chautauqua County, New York, United States. The lake is approximately long and wide at its greatest width. The surface area is approximately . The maximum depth is about . The shoreline is about of wh ...
in western New York, north into
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, throughout most of the
St Lawrence River The St. Lawrence River (french: Fleuve Saint-Laurent, ) is a large river in the middle latitudes of North America. Its headwaters begin flowing from Lake Ontario in a (roughly) northeasterly direction, into the Gulf of St. Lawrence, connecting ...
drainage, and northward throughout the upper
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
valley, although the species also extends as far south as
Chattanooga Chattanooga ( ) is a city in and the county seat of Hamilton County, Tennessee, United States. Located along the Tennessee River bordering Georgia, it also extends into Marion County on its western end. With a population of 181,099 in 2020, ...
in the
Tennessee River The Tennessee River is the largest tributary of the Ohio River. It is approximately long and is located in the southeastern United States in the Tennessee Valley. The river was once popularly known as the Cherokee River, among other names, ...
valley. Also, a small population is found in the Broad River in
South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...
. Several North Georgia reservoirs also have healthy stocked populations of muskie. They are also found in the Red River drainage of the
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
basin. Muskie were introduced to western Saint John River in the late 1960s and have now spread to many connecting waterways in northern
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and north ...
. The
Pineview Reservoir Pineview Dam is located in Ogden Canyon, east of Ogden, Utah, United States. Description Pineview Dam is located at the eastern end of Ogden Canyon at the confluence of the north, south, and center forks of the Ogden River. The western half o ...
in Utah is one of three Utah locations where the hybrid
Tiger Muskellunge The tiger muskellunge (''Esox masquinongy × lucius'' or ''Esox lucius × masquinongy''), commonly called tiger muskie, is a carnivorous fish, and is the usually-sterile, hybrid offspring of the true muskellunge (''Esox masquinongy'') and the no ...
is found. They prefer clear waters where they lurk along weed edges, rock outcrops, or other structures to rest. A fish forms two distinct home ranges in summer: a shallow range and a deeper one. The shallow range is generally much smaller than the deeper range due to shallow water heating up. A muskie continually patrols the ranges in search of available food in the appropriate conditions of water temperature.


Diet

Muskies are ambush predators who will swiftly bite their prey and then swallow it head first. Muskellunge are the top predator in any body of water where they occur and they will eat larger prey than most other freshwater fish. They eat all varieties of fish present in their ecosystem (including other muskellunge), along with the occasional
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
,
muskrat The muskrat (''Ondatra zibethicus'') is a medium-sized semiaquatic rodent native to North America and an introduced species in parts of Europe, Asia, and South America. The muskrat is found in wetlands over a wide range of climates and habitat ...
, rat,
mouse A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
,
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
, or
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form t ...
. They are capable of taking prey up to two-thirds of their body length due to their large stomachs. There have even been reports of large muskellunge attacking small
dogs The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Do ...
and even humans, although most of these reports are greatly exaggerated.


Length and weight

As muskellunge grow longer they increase in weight, but the relationship between length and weight is not linear. The relationship between them can be expressed by a power-law equation: :W = cL^b\!\, The exponent b is close to 3.0 for all species, and c is a constant for each species. For muskellunge, b = 3.325, higher than for many common species, and c = 0.000089 pounds/inch³. According to the
International Game Fish Association The International Game Fish Association (''IGFA'') is the leading authority on angling pursuits and the keeper of the most current World Record fishing catches by fish categories. Fishermen who are sport fishers are careful to follow their string ...
(IGFA) the largest muskellunge on record was
caught Caught is a method of dismissing a batsman in cricket. A batsman is out caught if the batsman hits the ball, from a legitimate delivery, with the bat, and the ball is caught by the bowler or a fielder before it hits the ground. If the ball h ...
by Cal Johnson in
Lac Courte Oreilles Lac Courte Oreilles ( ) is a large freshwater lake located in northwest Wisconsin in Sawyer County in townships 39 and 40 north, ranges 8 and 9 west. It is irregular in shape, having numerous peninsulas and bays, and is approximately six miles l ...
(recognized as Lake Courte Oreilles by the association), Hayward,
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
, on July 24, 1949. The
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
weighed 67 lbs 8 oz (30.61 kg) and was 60.25 in (153.035 cm) in length, and 33–34 in (83.82-86.36 cm) in girth.


Behavior

Muskellunge are sometimes
gregarious Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...
, forming small
schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...
in distinct territories. They spawn in mid to late spring, somewhat later than northern pike, over shallow, vegetated areas. A rock or sand bottom is preferred for spawning so the eggs do not sink into the mud and suffocate. The males arrive first and attempt to establish dominance over a territory. Spawning may last from five to 10 days and occurs mainly at night. The eggs are negatively buoyant and slightly adhesive; they adhere to plants and the bottom of the lake. Soon afterward, they are abandoned by the adults. Those
embryo An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male spe ...
s which are not eaten by fish, insects, or crayfish hatch within two weeks. The larvae live on
yolk Among animals which produce eggs, the yolk (; also known as the vitellus) is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example bec ...
until the mouth is fully developed, when they begin to feed on
copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...
s and other
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
. They soon begin to prey upon fish. Juveniles generally attain a length of by November of their first year.


Predators

Adult muskellunge are
apex predator An apex predator, also known as a top predator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the highest trophic lev ...
s where they occur naturally. Only humans and (rarely) large birds of prey such as
bald eagle The bald eagle (''Haliaeetus leucocephalus'') is a bird of prey found in North America. A sea eagle, it has two known subspecies and forms a species pair with the white-tailed eagle (''Haliaeetus albicilla''), which occupies the same niche as ...
s pose a threat to an adult but juveniles are consumed by other muskies, northern pike, bass,
trout Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-salmoni ...
, and occasionally birds of prey. The muskellunge's low reproductive rate and slow growth render populations highly vulnerable to
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in th ...
. This has prompted some jurisdictions to institute
artificial propagation Plant propagation is the process by which new plants grow from a variety of sources: seeds, cuttings, and other plant parts. Plant propagation can also refer to the man-made or natural dispersal of seeds. Propagation typically occurs as a step i ...
programs in an attempt to maintain otherwise unsustainably high rates of angling effort and
habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
.


Subspecies and hybrids

Though interbreeding with other pike species can complicate the classification of some individuals,
zoologists This is a list of notable zoologists who have published names of new taxa under the International Code of Zoological Nomenclature. A * Abe – Tokiharu Abe (1911–1996) * Abeille de Perrin, Ab. – Elzéar Abeille de Perrin (1843–1910) * ...
usually recognize up to three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of muskellunge. * The Great Lakes Muskellunge or Spotted Muskellunge (''E. m. masquinongy'') is the most common variety in the Great Lakes basin and surrounding area. The spots on the body form oblique rows. * The Chautauqua Muskellunge or Barred Muskellunge (''E. m. ohioensis'') is known from the Ohio River system,
Chautauqua Lake Chautauqua Lake is located entirely within Chautauqua County, New York, United States. The lake is approximately long and wide at its greatest width. The surface area is approximately . The maximum depth is about . The shoreline is about of wh ...
,
Lake Ontario Lake Ontario is one of the five Great Lakes of North America. It is bounded on the north, west, and southwest by the Canadian province of Ontario, and on the south and east by the U.S. state of New York. The Canada–United States border sp ...
, and the St Lawrence River. * The Clear Muskellunge (''E. m. immaculatus'') is most common in the inland lakes of Wisconsin, Minnesota, northwestern Ontario, and southeastern
Manitoba Manitoba ( ) is a Provinces and territories of Canada, province of Canada at the Centre of Canada, longitudinal centre of the country. It is Canada's Population of Canada by province and territory, fifth-most populous province, with a population o ...
. The
tiger muskellunge The tiger muskellunge (''Esox masquinongy × lucius'' or ''Esox lucius × masquinongy''), commonly called tiger muskie, is a carnivorous fish, and is the usually-sterile, hybrid offspring of the true muskellunge (''Esox masquinongy'') and the no ...
(''E. masquinongy'' × ''lucius'' or ''E. lucius'' × ''masquinongy'') is a
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
of the muskie and northern pike. Hybrids are sterile, although females sometimes unsuccessfully engage in spawning motions. Some hybrids are artificially produced and planted for anglers to catch. Tiger muskies grow faster than pure muskies, but do not attain the ultimate size of their pure relatives, as the tiger muskie does not live as long.


Attacks on humans

Although very rare, muskie attacks on humans do occur on occasion.


References


External links

{{DEFAULTSORT:Muskellunge Apex predators Esox Freshwater fish of the United States Fish of Canada Fish of the Eastern United States Fish of the Great Lakes Fish described in 1824 Taxa named by Samuel L. Mitchill Symbols of Wisconsin