Mulleripicus Pulverulentus
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The great slaty woodpecker (''Mulleripicus pulverulentus'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the family
Picidae Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. M ...
. It is found across the
Indian Subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
and
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. A unique and basically unmistakable bird, it is the largest species of woodpecker that is certain to exist today. It is a fairly gregarious species.


Range

It is found in the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
and
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, ranging across
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
,
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
,
Brunei Brunei ( , ), formally Brunei Darussalam ( ms, Negara Brunei Darussalam, Jawi alphabet, Jawi: , ), is a country located on the north coast of the island of Borneo in Southeast Asia. Apart from its South China Sea coast, it is completely sur ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
,
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
,
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
, the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
,
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
and
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
. It is found in the
Greater Sundas The Greater Sunda Islands (Indonesian and Malay: ''Kepulauan Sunda Besar'') are four tropical islands situated within Indonesian Archipelago, in the Pacific Ocean. The islands, Borneo, Java, Sulawesi and Sumatra, are internationally recognised fo ...
, but it does not inhabit
Bali Bali () is a province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller neighbouring islands, notably Nusa Penida, Nusa Lembongan, and Nu ...
.


Habitat

This species prefers to inhabit areas of primary semi-open, moist
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
and
tropical evergreen forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discont ...
though can on occasion range into adjacent secondary forests, clearings with scattered tall trees and similar almost park-like areas but do not generally visit heavily disturbed areas. Locally, the great slaty woodpecker prefers sprawling stands of
dipterocarp Dipterocarpaceae is a family of 16 genera and about 695 known species of mainly tropical lowland rainforest trees. The family name, from the type genus ''Dipterocarpus'', is derived from Greek (''di'' = two, ''pteron'' = wing and ''karpos'' = fru ...
and
teak Teak (''Tectona grandis'') is a tropical hardwood tree species in the family Lamiaceae. It is a large, deciduous tree that occurs in mixed hardwood forests. ''Tectona grandis'' has small, fragrant white flowers arranged in dense clusters (panicl ...
trees. Also found in mature sal forests,
swamp forest Freshwater swamp forests, or flooded forests, are forests which are inundated with freshwater, either permanently or seasonally. They normally occur along the lower reaches of rivers and around freshwater lakes. Freshwater swamp forests are found ...
and mangroves with tall, mature trees. The species usually occurs below an elevation of , but also locally in montane areas of up to , occasionally ranging up to .


Description

With the probable extinctions of the
imperial woodpecker The imperial woodpecker (''Campephilus imperialis'') is a woodpecker species endemic to Mexico. If it is not extinct, it is the world's largest woodpecker species, at long. Researchers have discovered that the imperial woodpecker has slow climb ...
and the ivory-billed woodpecker, this species, at long and a weight of , stands as the largest woodpecker in the world.''CRC Handbook of Avian Body Masses'' by John B. Dunning Jr. (Editor). CRC Press (1992), . The average weight was claimed in one study as .Styring, A. R., & bin Hussin, M. Z. (2004). ''Foraging ecology of woodpeckers in lowland Malaysian rain forests''. Journal of Tropical Ecology, 20(5), 487-494. Among standard measurements, the wing chord is , the
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...
is , the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
is and the tarsus is . This unique-looking woodpecker has several obvious distinctive features: a very long, strong chisel-tipped bill, an elongated neck and a long tail. A slight crest maybe occasionally evident. This species plumage is almost entirely dark grey or blackish slate-grey overlaid with small white spots. The throat is paler grey and males have small red moustache. Normally, the nominate subspecies is the darkest, most slaty gray race. ''M. p. harterti'' has a more pale throat with a greater amount of whitish feather tips forming small spot and is slightly paler below than the nominate, sometimes appearing almost whitish on the belly. The size and structure readily distinguishes this bird from almost any other species, including other woodpeckers. Occasionally, at first glance, the great slaty woodpecker is mistaken for a
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
but, obviously, such a resemblance is slight at best. For a bird of such great size, the great slaty woodpecker has a weak, quiet voice, especially compared to other large woodpeckers, which tend to have loud, booming voices. The species call is a whinnying cackle of 2 to 5, usually 4 notes, ''woikwoikwoikwoik'', the initial being higher in pitched and the middle note being distinctly lower. Single ''dwot'' calls, variable in sound, strength and duration, are sometimes given while perched or in flight. Breeding pairs of these woodpeckers have been heard to softly mew at each other. In more antagonistic situations, sharp ''taw-whit'' or ''dew-it'' calls are uttered while the birds swing their heads back and forth.


Ecology

Great slaty woodpeckers are mostly seen in groups consisting of 3 to 6 individuals, which consist of a breeding pair and their young from prior years. Groups often forage on shared feeding sites in the form of nests of social insects as ants,
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
s, wood-boring
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s and stingless bees. Ants seem to be generally favored in the diet, though larvae of other species may be eaten quite regularly as well. Occasionally, small
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
may supplement the diet. Females spend more time searching for feeding sources and males, which have slightly larger bills, spend more time opening the sources. Preferred feeding sources are mostly found in large branches or trunks of large, living trees. The groups will travel considerable distance to access these trees and, as such, the home ranges of the species are quite large. Occasionally, though, they will feed at lower levels in trees and even amongst saplings. Usually, feeding groups of these woodpeckers do not linger in any given area for long. Sometimes this species associates with slightly smaller white-bellied woodpeckers and considerably smaller
greater flameback The greater flameback (''Chrysocolaptes guttacristatus'') also known as greater goldenback, large golden-backed woodpecker is a woodpecker species. It occurs widely in the northern Indian subcontinent, eastwards to southern China, the Malay Peni ...
s, with the foraging methods of the very different woodpeckers minimizing competition between the species. Perhaps more considerable competition for food sources generally comes in the form of
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
s and arboreal (or tree-dwelling) mammals. The great slaty woodpecker usually works a tree upwards and, though capable of swifter movements, has been described while foraging as if moving in "slow motion". It forages by gleaning, probing, pecking, prising off bark and hammering with powerful and loud blows to excavate the wood. Gleaning is the most important foraging method for the species, with the long neck and bill allowing it to reach out over a considerable distance into the cracks and crevices of trees. This species often flies high over the trees for long distances between successful foraging patches. In flight, its feather rustle noisily. The great slaty woodpecker usually engages in less dipping during than other woodpeckers and flies in a mixed flying style described as quite crow-like. Like all woodpeckers, breeding pairs roost in separate tree holes but regularly vocalized to stay in contact. The pair bond appears to be lifelong. These woodpeckers engage in displays, largely for territorial purposes. Displays include head-swinging, where the appears to lag behind the body in swinging movements, whinnying calls and widen their wings and tail considerably. Few nests of the species have been described in detail, but at least occasionally nests are raised cooperatively by groups. Known nests, at anywhere from in height in the trees, were located in very large trees. When excavating the nest hole, both parents participate but reportedly the male does the majority of the work. The nest hole entrance will be around across, but much wider inside the tree. The pair will only use a nest from a prior year if competition is too overbearing for a newly constructed hole. The nesting season, in
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
at least, appears to be from March to August. The clutch reportedly consists of two to four eggs, which are incubated by both parents. Both parents also feed and generally brood the young. The young great slaty woodpeckers probably stay with their parents until the next breeding season.


Status

Probably because of their feeding and breeding dependence on large old trees, great slaty woodpeckers are most common in primary forests and show density reductions of over 80% in logged forests. The global population is in decline because of the loss of forest cover and logging of old-growth forest throughout its range, with habitat loss being particularly rapid in Myanmar, Cambodia and Indonesia which are the countries that still hold the majority of the global population. In 2010, the great slaty woodpecker was included in the IUCN Red List in the Vulnerable category.


References


External links

* BirdLife International 2011. Species factsheet: Great slaty woodpecke

Downloaded on 1 November 2011.
Image at ADW


Images at Arkive {{Taxonbar, from=Q1266445 Mulleripicus, great slaty woodpecker Birds of Southeast Asia Birds described in 1826 Taxonomy articles created by Polbot