Mullein Wave
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''Scopula marginepunctata'', the mullein wave, is a
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
of the family
Geometridae The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek ''geo'' γεω (derivative form of or "the earth"), and ''metro ...
. It was described by
Johann August Ephraim Goeze Johann August Ephraim Goeze (; 28 May 1731 – 27 June 1793) was a German zoologist, born in Aschersleben. He is known for the discovery of tardigrades, also called water bears. He was the son of Johann Heinrich and Catherine Margarete (née Kirc ...
in 1781. It is found throughout Europe.


Distribution

''Scopula marginepunctata'' occurs in Europe from the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
in the west to the
Ural mountains The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western ...
in the east. In the north, the range extends to the south coast of
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
, the southern
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
and the German
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and ...
coast. However the species is missing in parts of northern Germany. There are isolated occurrences on Bornholm, Skåne and the southern Baltic. In the south, the range extends in North Africa from
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
in the west to
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Medit ...
. The distribution ranges from there further over the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabia (including the Arabian Peninsula and Bahrain), Asia Minor (Asian part of Turkey except Hatay Province), East Thrace (Europ ...
,
Asia minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
, the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia (country), Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range ...
, northern
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...
and
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
.


Description

The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...
is . The length of the forewings is . The second generation is often smaller and the moths reach only about 18 mm wingspan. Colour and wing pattern vary. The base colour is off-white to light brown, and the wings are strong dark over dusted with dark scales. In the Swiss Canton of Ticino darkened forms occur, which were formerly called subspecies ''insubrica''. The pattern of the wing is dark grey to dark brown, depending on the ground colour. The interior cross line and median bands are usually little developed. The medium band is relatively wide, if present, but frequently washed out. The interior cross line is often reduced to a row of dots. Usually only the outer cross line is significantly developed and almost always very pronounced and jagged, the tips of the spikes are even darker highlighted towards the marginal field. The front edge itself may be slightly darker than the ground colour. In the marginal field, usually four to five tooth-shaped, slurred stains are present, two penultimate often characteristic close. The intensity of each cross line may be slightly different on forewings and hindwings. Marginal stains are usually present. Discal flecks are almost always present on the front as well as on the rear wings. In North Africa the subspecies ''Scopula marginepunctata argillacea'' replaces the nominate. It is light brown with relatively weak pattern. In the north of Iran, Central Asia and Mongolia, the subspecies ''Scopula marginepunctata terrigena'' occurs. It is slightly larger than the nominate with a wingspan. The marginal field and medium fields are wide and slightly dark brown in colour. The egg is elongated oval and flattened at the top. It is initially coloured light yellow, just before hatching it turns stained red. The outside shows strong longitudinal ribs that intersect with slightly weaker cross ribs. The caterpillar is relatively long and slender and gets slightly thinner forward. The segments are clearly constricted. It is grey-yellow to grey-brown coloured with a dark dorsal line. The dorsal line is accompanied by black dots. The abdominal side is slightly darker, the spiracles are coloured black. The pupa is light brown coloured with slightly greenish wing sheaths. The head and posterior end are a little darker. The segments are relatively deep, the cremaster is relatively short and studded with bristles.


Biology

The moth flies in two generations from mid-May to September and the larva feeds on
yarrow ''Achillea millefolium'', commonly known as yarrow () or common yarrow, is a flowering plant in the family Asteraceae. Other common names include old man's pepper, devil's nettle, sanguinary, milfoil, soldier's woundwort, and thousand seal. The ...
and
mugwort Mugwort is a common name for several species of aromatic flowering plants in the genus '' Artemisia.'' In Europe, mugwort most often refers to the species ''Artemisia vulgaris'', or common mugwort. In East Asia the species ''Artemisia argyi'' is ...
.


Subspecies

*''Scopula marginepunctata marginepunctata'' *''Scopula marginepunctata subatrata'' (Wagner, 1919) *''Scopula marginepunctata terrigena'' Prout, 1935


References


External links


Mullein wave on UKMothsLepiforum.deVlindernet.nl
{{Taxonbar, from=Q143162 marginepunctata Moths described in 1781 Moths of Africa Moths of Asia Moths of Europe Taxa named by Johann August Ephraim Goeze