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The mouflon (''Ovis gmelini'') is a
wild sheep ''Ovis'' is a genus of mammals, part of the Caprinae subfamily of the ruminant family Bovidae. Its seven highly sociable species are known as sheep or ovines. Domestic sheep are members of the genus, and are thought to be descended from the wil ...
native to
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
, the
Caspian Caspian can refer to: *The Caspian Sea *The Caspian Depression, surrounding the northern part of the Caspian Sea *The Caspians, the ancient people living near the Caspian Sea *Caspian languages, collection of languages and dialects of Caspian peopl ...
region from eastern Turkey, Armenia, Azerbaijan, and Iran. It is thought to be the ancestor of all modern
domestic sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated sh ...
breeds.


Taxonomy

''Ovis gmelini'' was the
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
proposed by
Edward Blyth Edward Blyth (23 December 1810 – 27 December 1873) was an English zoologist who worked for most of his life in India as a curator of zoology at the museum of the Asiatic Society of India in Calcutta. Blyth was born in London in 1810. In 1841 ...
in 1841 for wild sheep in the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
. In the 19th and 20th centuries, several wild sheep were described that are considered mouflon
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
today: * ''Ovis ophion'' by Blyth in 1841 for wild sheep in
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
; * ''Ovis laristanica'' by Nikolai Nasonov in 1909 for wild sheep in Lar in southern Iran; * ''Ovis orientalis isphahanica'' by Nasonov in 1910 for wild sheep in the
Zagros Mountains The Zagros Mountains ( ar, جبال زاغروس, translit=Jibal Zaghrus; fa, کوه‌های زاگرس, Kuh hā-ye Zāgros; ku, چیاکانی زاگرۆس, translit=Çiyakani Zagros; Turkish: ''Zagros Dağları''; Luri: ''Kuh hā-ye Zāgro ...
.


Subspecies

Five mouflon
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of are distinguished by
MSW3 ''Mammal Species of the World: A Taxonomic and Geographic Reference'' is a standard reference work in mammalogy giving descriptions and bibliographic data for the known species of mammals. It is now in its third edition, published in late 2005 ...
: *
Armenian mouflon The Armenian mouflon (''Ovis gmelini gmelini'') is an endangered subspecies of mouflon endemic to Iran, Armenia, Nakhchivan Autonomous Republic of Azerbaijan, Turkey and Iraq. Taxonomy The Armenian mouflon was first described in 1840 by Edward Bl ...
(Armenian red sheep), ''O. g. gmelini'' (Blyth, 1851):
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
; native to northwestern
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ''Ox ...
, and
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...
. It has been introduced to Texas in the U.S. *
Esfahan mouflon Isfahan ( fa, اصفهان, Esfahân ), from its Achaemenid empire, ancient designation ''Aspadana'' and, later, ''Spahan'' in Sassanian Empire, middle Persian, rendered in English as ''Ispahan'', is a major city in the Greater Isfahan Regio ...
, ''O. g. isphahanica'' (Nasonov, 1910):
Zagros Mountains The Zagros Mountains ( ar, جبال زاغروس, translit=Jibal Zaghrus; fa, کوه‌های زاگرس, Kuh hā-ye Zāgros; ku, چیاکانی زاگرۆس, translit=Çiyakani Zagros; Turkish: ''Zagros Dağları''; Luri: ''Kuh hā-ye Zāgro ...
, Iran. * Laristan mouflon, ''O. g. laristanica'' (Nasonov, 1909): a small subspecies, its range is restricted to some desert reserves near Lar in southern Iran. * Cyprus mouflon, ''O. g. ophion'' (Blyth, 1841): also called ''agrino'' (from the Greek ); nearly driven to extinction during the 20th century. In 1997, about 1,200 individuals were counted. The television show ''
Born to Explore with Richard Wiese Richard Wiese (born July 13, 1959) is an American explorer, the longest serving President of The Explorers Club, and Executive Producer and Host of the multiple Emmy Award-winning ABC and PBS program, Born to Explore. Early life and education ...
'' reported 3,000 individuals on
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
. The
European mouflon The European mouflon (''Ovis aries musimon'') is a feral subspecies of the primitive domestic sheep. It was originally found only on the Mediterranean islands of Corsica and Sardinia, but has since been introduced into many other regions o ...
was once thought to be a subspecies of the mouflon, but is now considered to be a feral descendant of the domestic sheep (''Ovis aries''), as ''Ovis aries musimon''.


Relation to other sheep

Based on comparison of mitochondrial
cytochrome b Cytochrome b within both molecular and cell biology, is a protein found in the mitochondria of eukaryotic cells. It functions as part of the electron transport chain and is the main subunit of transmembrane cytochrome bc1 and b6f complexes. F ...
gene sequences, three groups of sheep (''
Ovis ''Ovis'' is a genus of mammals, part of the Caprinae subfamily of the ruminant family Bovidae. Its seven highly sociable species are known as sheep or ovines. Domestic sheep are members of the genus, and are thought to be descended from the wil ...
'') have been identified: Pachyceriforms of Siberia (
snow sheep The snow sheep (''Ovis nivicola''), or Siberian bighorn sheep, is a species of sheep from the mountainous areas in the northeast of Siberia. One subspecies, the Putorana snow sheep ''(Ovis nivicola borealis)'', lives isolated from the other forms ...
) and North America (
bighorn The bighorn sheep (''Ovis canadensis'') is a species of sheep native to North America. It is named for its large horns. A pair of horns might weigh up to ; the sheep typically weigh up to . Recent genetic testing indicates three distinct subspec ...
and Dall sheep), Argaliforms (
argali The argali (''Ovis ammon''), also known as the mountain sheep, is a wild sheep that roams the highlands of western East Asia, the Himalayas, Tibet, and the Altai Mountains. Description The name 'argali' is the Mongolian word for wild sheep. It ...
) of Central Asia, and Moufloniforms (
urial The urial ( ; ''Ovis vignei''), also known as the arkars or shapo, is a wild sheep native to Central and South Asia. It is listed as Vulnerable on the IUCN Red List. Characteristics Urial males have large horns, curling outwards from the to ...
, mouflon, and
domestic sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated sh ...
) of Eurasia.Bunch, Wu, Zhang, Wang (2005). "Phylogenetic analysis of the snow sheep (''Ovis nivicola'') and closely related taxa", ''Journal of Heredity'', 97 (1) 21–30

/ref> However, a comparison of the
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
control region (CR) found that two subspecies of urial, ''Ovis vignei'' (or ''orientalis'') ''arkal'' and ''O. v./o. bochariensis'', grouped with two different clades of argali (''Ovis ammon''). The ancestral sheep is presumed to have had 60
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
s, as in goats (''Capra''). Mouflon and domestic sheep have 54 chromosomes, with three pairs (1+3, 2+8, 5+11) of ancestral acrocentric chromosomes joined to form bi-armed chromosomes. This is in contrast to the argali and urial, which have 56 and 58 chromosomes respectively. If the urial is as closely related to the mouflons as mitochondrial DNA indicates, then two chromosomes would need to have split during its evolution away from the mouflon (sub)species.


Description

Mouflon have reddish to dark brown, short-haired coats with dark back stripes and black ventral areas and light-colored saddle patches. The males are
horned A horn is a permanent pointed projection on the head of various animals that consists of a covering of keratin and other proteins surrounding a core of live bone. Horns are distinct from antlers, which are not permanent. In mammals, true horns ...
; some females are horned, while others are polled. The horns of mature rams are curved almost one full revolution (up to 85 cm). Mouflon have shoulder heights of around 0.9 m and body weights of 50 kg (males) and 35 kg (females).


Distribution and habitat

The mouflon occurs in the
Lesser Caucasus The Lesser Caucasus, also called Caucasus Minor, is the second of the two main mountain ranges of Caucasus mountains, of length about . The western portion of the Lesser Caucasus overlaps and converges with east Turkey and northwest Iran. It runs ...
in southeastern
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
,
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ''Ox ...
and
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...
, as well as in
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
's western
Alborz The Alborz ( fa, البرز) range, also spelled as Alburz, Elburz or Elborz, is a mountain range in northern Iran that stretches from the border of Azerbaijan along the western and entire southern coast of the Caspian Sea and finally runs nort ...
region and the
Zagros Mountains The Zagros Mountains ( ar, جبال زاغروس, translit=Jibal Zaghrus; fa, کوه‌های زاگرس, Kuh hā-ye Zāgros; ku, چیاکانی زاگرۆس, translit=Çiyakani Zagros; Turkish: ''Zagros Dağları''; Luri: ''Kuh hā-ye Zāgro ...
spanning across eastern
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
and western Iran. It was possibly introduced to
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
during the
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
period.


Behaviour and ecology


Reproduction

Mouflon rams have a strict dominance hierarchy. Before mating season or "rut", which is from late autumn to early winter, rams try to create a dominance hierarchy to determine access to ewes (female mouflon) for mating. Mouflon rams fight one another to obtain dominance and win an opportunity to mate with females. Mouflons reach sexual maturity at the age of 2 to 4 years. Young rams need to obtain dominance before they get a chance to mate, which takes another 3 years for them to start mating. Mouflon ewes also go through a similar hierarchy process in terms of social status in the first 2 years, but can breed even at low status. Pregnancy in females lasts 5 months, in which they produce one to two offspring. A mouflon was cloned successfully in early 2001, and lived at least seven months, making it the first clone of an endangered mammal to survive beyond infancy. This demonstrated that a common species (in this case, a domestic sheep) can successfully become a surrogate for the birth of an exotic animal such as the mouflon. If cloning of the mouflon can proceed successfully, it has the potential to reduce strain on the number of living specimens.


Conservation

The mouflon is protected in Armenia and Azerbaijan. In Turkey and Iran, hunting is only allowed with a special license. The population in Cyprus is listed as a strictly protected species in the
Habitats Directive The Habitats Directive (more formally known as Council Directive 92/43/EEC on the Conservation of natural habitats and of wild fauna and flora) is a directive adopted by the European Community in 1992 as a response to the Berne Convention. The E ...
of the European Union and has been listed in
CITES Appendix I CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
since November 2019.


In culture

The wild sheep of Corsica were locally called ''mufro'' (male) and ''mufra'' (female). The French naturalist Buffon (1707–1788) rendered this in French as '. The male mouflon is denominated in Corsica ''Mufro'', and the female ''Mufra'', from which the word ''Moufflon'' was formed; in Sardinia, the male is called ''Murvoni'', and the female ''Murva'', though it is not unusual to hear the peasants style both indiscriminately ''Mufion'', which is a palpable corruption of the Greek ''Ophion''. * The mouflon is featured on the symbol of
Cyprus Airways Cyprus Airways (Greek: Κυπριακές Αερογραμμές) is the flag carrier airline of Cyprus, based at Larnaca International Airport. It commenced operations on 1 June 2017. History Cyprus Airways resumed operations in 2016 after win ...
, as well as on the 1-, 2-, and 5-cent
Cypriot euro coins Cypriot euro coins feature three separate designs for the three series of coins. Cyprus has been a member of the European Union since 1 May 2004, and is a member of the Economic and Monetary Union of the European Union. It has completed the third ...
. * The mouflon is featured on the historical flag of the
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ''Ox ...
n
kingdom Kingdom commonly refers to: * A monarchy ruled by a king or queen * Kingdom (biology), a category in biological taxonomy Kingdom may also refer to: Arts and media Television * ''Kingdom'' (British TV series), a 2007 British television drama s ...
of Syunik, as well as on tombstones. * The mouflon is featured both on the symbol and as the nickname of the
Cyprus national rugby union team The Cyprus national rugby union team is nicknamed ‘’The Mighty Mouflons’’ after a kind of horned sheep which is also the republic's national animal. They have won 28 of their 30 games and hold the world record for the most consecutive i ...
.


See also

* Castlemilk Moorit *
Urial The urial ( ; ''Ovis vignei''), also known as the arkars or shapo, is a wild sheep native to Central and South Asia. It is listed as Vulnerable on the IUCN Red List. Characteristics Urial males have large horns, curling outwards from the to ...


References

{{Authority control Ovis Mammals of Azerbaijan Mammals of the Middle East Mammals of Turkey Caucasus Fauna of Iran Mammals described in 1841 Taxa named by Edward Blyth