The mottled umber (''Erannis defoliaria'') is a

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

of the family

Geometridae The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek ''geo'' γεω (derivative form of or "the earth"), and ''met ...

. It is common throughout much of the

Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...

region. The species was first described by

Carl Alexander Clerck Carl Alexander Clerck (1709 – 22 July 1765) was a Sweden, Swedish entomologist and arachnology, arachnologist. Clerck came from a family in the petty Swedish nobility, nobility and entered the University of Uppsala in 1726. Little is known of hi ...

in 1759.

Distribution

The species can be found in western Europe from northern

Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...

to the

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...

, then east to the

Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...

, northern

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admini ...

, and

Ussuri The Ussuri or Wusuli (russian: Уссури; ) is a river that runs through Khabarovsk and Primorsky Krais, Russia and the southeast region of Northeast China. It rises in the Sikhote-Alin mountain range, flowing north and forming part of the Si ...

Description

The female of this species is wingless and rather

spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...

-like and can be found on the trunks and stems of its

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...

l food

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...

s. She is white or yellow-and-black patched. The male is fully winged (

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...

40–45 mm) and very variable. The ground colour of the forewing is pale yellow or light yellow ochre and suffused dark grey. The basal and distal fasciae are dark brown. The basal fascia is bordered on the inner side by a brown cloud; the distal fascia has a brown cloud on the outer edge. There is also a brown cloud along the costa. There are two dark brown oblong spots in the upper half of the terminal area. The hindwing is grey white with grey suffusion. There is a grey-black discal spot on both wings. There are numerous and significant deviations from this colour pattern and some specimens are very dark. See Prout (1912–16) Truly melanic forms are also frequent. As with the

peppered moth The peppered moth (''Biston betularia'') is a temperate species of night-flying moth. It is mostly found in the northern hemisphere in places like Asia, Europe and North America. Peppered moth evolution is an example of population genetics an ...

, the darker forms tend to be prevalent in industrial areas.

Biology

The adults have been recorded from September to December in the Great Britain and Ireland. Males are attracted to light. The egg is oval with very shallow, somewhat zigzag longitudinal ridges, a cell-pattern only discernible on high magnification and brownish yellow, without gloss. The larva is also variable from cream to dark brown and quite gaily coloured. It feeds on a wide variety of

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s and

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

s (see list below) and can be a serious pest in

orchard An orchard is an intentional plantation of trees or shrubs that is maintained for food production. Orchards comprise fruit- or nut-producing trees which are generally grown for commercial production. Orchards are also sometimes a feature of larg ...

s. The species overwinters as an

egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...

. ; Larval food plants include, * apple (''

Malus ''Malus'' ( or ) is a genus of about 30–55 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples, wild apples, and rainberries. The genus is native to the temperate zone of th ...

'' species) * birch – silver (''

Betula pendula ''Betula pendula'', commonly known as silver birch, warty birch, European white birch, or East Asian white birch, is a species of tree in the family Betulaceae, native to Europe and parts of Asia, though in southern Europe, it is only found a ...

''), downy (''

Betula pubescens ''Betula pubescens'' (syn. ''Betula alba''), commonly known as downy birch and also as moor birch, white birch, European white birch or hairy birch, is a species of deciduous tree, native and abundant throughout northern Europe and northern Asia ...

'') * blackthorn (''

Prunus spinosa ''Prunus spinosa'', called blackthorn or sloe, is a species of flowering plant in the rose family Rosaceae. The species is native to Europe, western Asia, and regionally in northwest Africa. It is locally naturalized in New Zealand, Tasmania, ...

'') * dog rose (''

Rosa canina ''Rosa canina'', commonly known as the dog rose, is a variable climbing, wild rose species native to Europe, northwest Africa, and western Asia. Description The dog rose is a deciduous shrub normally ranging in height from , though sometimes it ...

'') * elm (''

Ulmus Elms are deciduous and semi-deciduous trees comprising the flowering plant genus ''Ulmus'' in the plant family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of Nor ...

'' species) * field maple (''

Acer campestre ''Acer campestre'', known as the field maple, is a flowering plant species in the family Sapindaceae. It is native to much of continental Europe, Britain, southwest Asia from Turkey to the Caucasus, and north Africa in the Atlas Mountains. It has ...

'') * hawthorn – midland (''

Crataegus laevigata ''Crataegus laevigata'', known as the Midland hawthorn, English hawthorn, woodland hawthorn, or mayflower, is a species of hawthorn native to western and central Europe, from Great Britain (where it is typically found in ancient woodland and old ...

'') common (''

Crataegus monogyna ''Crataegus monogyna'', known as common hawthorn, one-seed hawthorn, or single-seeded hawthorn, is a species of flowering plant in the rose family Rosaceae. It is native to Europe, northwestern Africa, and West Asia, but has been introduced in ...

'') * hazel (''

Corylus avellana ''Corylus avellana'', the common hazel, is a species of flowering plant in the birch family Betulaceae. It is native to Europe and western Asia. It is an important component of the hedgerows that were the traditional field boundaries in lowland En ...

'') * hornbeam ('' Carpinus betulus'') * oak – sessile (''

Quercus petraea ''Quercus petraea'', commonly known as the sessile oak, Cornish oak, Irish Oak or durmast oak, is a species of oak tree native to most of Europe and into Anatolia and Iran. The sessile oak is the national tree of Ireland, and an unofficial emble ...

''), pedunculate (''

Quercus robur ''Quercus robur'', commonly known as common oak, pedunculate oak, European oak or English oak, is a species of flowering plant in the beech and oak family, Fagaceae. It is a large tree, native plant, native to most of Europe west of the Caucasus ...

'') * sallow – (''

Salix Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist so ...

'' species) including goat (''

Salix caprea ''Salix caprea'', known as goat willow, pussy willow or great sallow, is a common species of willow native to Europe and western and central Asia.Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook 4. . Des ...

'') * Sitka spruce (''

Picea sitchensis ''Picea sitchensis'', the Sitka spruce, is a large, coniferous, evergreen tree growing to almost tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth-larg ...

'') * sweet chestnut (''

Castanea sativa ''Castanea sativa'', the sweet chestnut, Spanish chestnut or just chestnut, is a species of tree in the family Fagaceae, native to Southern Europe and Asia Minor, and widely cultivated throughout the temperate world. A substantial, long-lived ...

'') * sycamore (''

Acer pseudoplatanus ''Acer pseudoplatanus'', known as the sycamore in the British Isles and as the sycamore maple in the United States, is a species of flowering plant in the Sapindus, soapberry and lychee family Sapindaceae. It is a large deciduous, broad-leaved ...

'')

Similar species

*'' Agriopis aurantiaria'' *''

Agriopis marginaria The dotted border (''Agriopis marginaria'') is a moth of the family Geometridae. The species was first described by Johan Christian Fabricius in 1776. It is found throughout Europe, except the far north, and the Near East. The female of this ...

Gallery

Image:Erannis defoliaria larve.jpg, Caterpillar File:Erannis defoliaria pupa.JPG, Pupa Image:Erannis defoliaria female.jpg, Wingless female Image:Britishentomologyvolume6Plate703.jpg, Illustration from John Curtis's ''

British Entomology ''British Entomology'' is a classic work of entomology by John Curtis, FLS. It is subtitled ''Being Illustrations and Descriptions of the Genera of Insects found in Great Britain and Ireland: Containing Coloured Figures from Nature of the Most ...

'' Volume 6

References

* *

External links

* * {{Taxonbar , from=Q1440936 Bistonini Moths described in 1759 Moths of Asia Moths of Europe Moths of Iceland Moths of Japan Taxa named by Carl Alexander Clerck