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''Morpho anaxibia'', the Anaxibia morpho, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
Neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
.


Description

In 1913,
Hans Fruhstorfer Hans Fruhstorfer (7 March 1866, in Passau, Germany – 9 April 1922, in Munich) was a German explorer, insect trader and entomologist who specialised in Lepidoptera. He collected and described new species of exotic butterflies, especially in Ada ...
wrote: "''M. anaxibia'' Esp. (69 c) may be regarded as one of the characteristic butterflies of southern Brazil. The species stands quite alone in having the abdomen blue above, but forms a transition from the '' rhetenor'' to the ''
menelaus In Greek mythology, Menelaus (; grc-gre, Μενέλαος , 'wrath of the people', ) was a king of Mycenaean (pre- Dorian) Sparta. According to the ''Iliad'', Menelaus was a central figure in the Trojan War, leading the Spartan contingent of th ...
'' group, ''anaxibia'', however, lacks the wonderful gloss of the ''rhetenor'' series, the blue is duller, although it has its own particular beauty, and the dazzling iridescence is wanting. The male still suggests ''rhetenor'' in the under surface, the female reveals more connection with the ''menelaus'' group, but its marginal spots are yellow instead of white. The male bears a narrow black costal border, which is inappreciably widened at the apex and then continued to the hinder angle of the forewing as a proximally more or less dentate distal border. Beyond the cell there is a white patch at the costal margin, but in examples from
Blumenau Blumenau is a city in Vale do Itajaí, state of Santa Catarina (Brazil), Santa Catarina, in the South Region, Brazil, South Region of Brazil. It is away from the state capital of Florianópolis. The city was founded by the German chemist an ...
this is frequently absent. The forewing is predominantly black beneath, the hindwing red-brown, respectively with two to three and four to five medium-sized eye-spots, slightly pupilled with white and mostly ringed with light grey-yellow. The under surface of the female is more variegated, marked with slightly glossy broad grey zigzag bands and patches. The female varies above in the white, delicately blue-dusted transcellular patch of the forewing, which may be composed of only one (Parana) or of three divisions."Fruhstorfer, H., 1913. Family: Morphidae. In A. Seitz (editor), ''Macrolepidoptera of the World'', vol. 5: 333–356. Stuttgart: Alfred Kernen. — Larva on ''Canella'' and one of the Myrtaceae "Grumexama", elongate, appreciably thickened in the middle, head with two lateral, not very distinctly projecting tubercles with long bristles. Head shining yellowish horn-colour, covered with small rounded pitting and dots, with fine white hairs and with strong red-brown bristles. Body yellow, sides densely clothed with fine woolly hairs, back gaily coloured, the first two segments finely haired, on the third to the sixth segment long lateral tufts of bristles, then a varied mixture of black, bordeaux-red and white. The 5th and 6th segments naked, the 7th and 8th with a St. Andrew's cross-shaped figure, the four ends of which are adorned with gay tufts of bristles. Anus with two lateral bunches of bristles and the beginning of the last segment, like the anterior ones, armed with brightly coloured hairs. Over the back runs a rather broad band with the inner edge, distinct, the outer broken up into marbling. All the segments further with a double red lateral line. Underside with red-brown patches between the legs. Pupa posteriorly strongly swollen, short, light green with yellowish wing-cases and sharply defined yellow-white ring shortly behind the thickest part of the body. Head with two fine, pointed black horns ; in general smaller, weaker, slighter than those of Morpho hercules Dalm. In the yellowish ring the pupa of anaxibia also differs from that of ''M. catenari'' MS Perty. In Sta. Catharina pupation takes place approximately from the 15th of January to the 1st of March."


Behaviour

"According to Mabille it flies for 15–30 days at the end of January and beginning of February and the time of flight only occasionally extends to the end of March. I myself observed ''anaxibia'' in large numbers in Santa Catharina, in February, particiuarly in a side valley of the Capivary River. Here a crystal waterfall sprinkled the roots of the forest giants and thus provided a centre of attraction for butterflies" of all sorts. As a wild beast seeks out certain set tracks in order to reach the water, so ''anaxibia'' also haunts this spot daily, following up accidental clearings in order to gain the cascade, as if they also sought there for cooling refreshment among the softly rustling arches of the tall bamboos shaken and bedewed by the foaming water. They did not arrive in companies but singly, sailing quietly along, but nevertheless after heavy rain the sand-banks before the rocks of the river were strewed with the glittering blue wings of dead ''anaxibia'' and ''aegea'' which the water washed up. The female is extraordinarily rare and on hot days rests in the morning hours with the wings closed in wet places in the forest-paths where it can imbibe the moisture. It is then so well concealed by the red under surface, which differs but little from the surrounding ground, that it is usually only noticed by the collector when it is too late. According to von Bonninghausen ''anaxibia'' is common in March on the Coreovado."


Subspecies

Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
have been described: * ''Morpho anaxibia pelias'' Fruhstorfer, 1913"''Morpho'' Fabricius, 1807"
at Markku Savela's ''Lepidoptera and Some Other Life Forms''
* ''Morpho anaxibia
calliphon Calliphon (or Callipho, grc-x-koine, Καλλιφῶν) was a Greek philosopher, who probably belonged to the Peripatetic school and lived in the 2nd century . He is mentioned several times and condemned by Cicero Marcus Tullius Cicero ( ; ...
'' Fruhstorfer, 1916


Larval food plants

Larvae have been recorded feeding on a wide range of plants:
Canellaceae The Canellaceae are a family of flowering plants in the order Canellales.Walter S. Judd, Christopher S. Campbell, Elizabeth A. Kellogg, Peter F. Stevens, and Michael J. Donoghue. 2008. ''Plant Systematics: A Phylogenetic Approach'', Third Editio ...
,
Guttiferae The Clusiaceae or Guttiferae Juss. (1789) (''nom. alt. et cons.'' = alternative and valid name) are a family of plants including 13 genera and ca 750 species. Several former members of Clusiacae are now placed in Calophyllaceae and Hypericacea ...
,
Erythroxylaceae Erythroxylaceae (the coca family) is a family of flowering trees and shrubs consisting of 4 genera and 271 species. The four genera are ''Aneulophus'' Benth., ''Erythroxylum'' P.Browne, ''Nectaropetalum'' Engl., and ''Pinacopodium'' Exell & Mendo ...
,
Myrtaceae Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All speci ...
,
Moraceae The Moraceae — often called the mulberry family or fig family — are a family of flowering plants comprising about 38 genera and over 1100 species. Most are widespread in tropical and subtropical regions, less so in temperate climates; however ...
and
Lauraceae Lauraceae, or the laurels, is a plant family that includes the true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genera worldwide (Christenhusz & Byng 2016 ). They are dicotyledons, and occur ma ...
.


Etymology

Anaxibia Anaxibia (; Ancient Greek: ) is the name of six characters in Greek mythology. *Anaxibia, one of the Danaïdes, married to Archelaus, son of Aegyptus. *Anaxibia, a naiad of the Ganges river. She fled from the advances of Helios, but she disapp ...
was the daughter of Bias and Iphianassa in Greek mythology. She married
Pelias Pelias ( ; Ancient Greek: Πελίας) was king of Iolcus in Greek mythology. He was the one who sent Jason on the quest for the Golden Fleece. Family Pelias was the son of Tyro and Poseidon. His wife is recorded as either Anaxibia, daughter ...
, the name given to one of the two subspecies.


References

* Le Moult (E.) & Réal (P.), 1962–1963. ''Les Morpho d'Amérique du Sud et Centrale'', Editions du cabinet entomologique E. Le Moult, Paris. *Paul Smart, 1976 ''The Illustrated Encyclopedia of the Butterfly World in Color''. London, Salamander: ''Encyclopedie des papillons''. Lausanne, Elsevier Sequoia (French language edition) page 233 fig.11, female (Brazil).


External links


Butterflies of America
Images of type and other specimens. {{Taxonbar, from=Q2412551 Morpho Nymphalidae of South America Endemic fauna of Brazil Fauna of Brazil