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Morelet's crocodile ''(Crocodylus moreletii)'', also known as the Mexican crocodile or Belize crocodile, is a modest-sized
crocodilian Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
found only in fresh waters of the Atlantic regions of
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
,
Belize Belize (; bzj, Bileez) is a Caribbean and Central American country on the northeastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a wate ...
and
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
. It usually grows to about in length. It is a
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
species. The species has a fossil record in
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
.


Taxonomy and etymology

Morelet's crocodile was first described in 1850 in Mexico by the French naturalist
Pierre Marie Arthur Morelet Pierre Marie Arthur Morelet (26 August 1809 – 9 October 1892) was a French natural history, naturalist, born in Lays, Doubs. He was a member of the Commission to Algeria, primarily as a natural artist, drawing any natural findings. He collect ...
. The species was subsequently named after him. It was long confused with the
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
and
Cuban crocodile The Cuban crocodile (''Crocodylus rhombifer'') is a small-medium species of crocodile endemic to Cuba. Typical length is and typical weight . Large males can reach as much as in length and weigh more than . Despite its smaller size, it is a hig ...
s because of similar characteristics and an ambiguous type locality. It was not generally accepted as a separate species until the 1920s.


Evolution

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
'' Crocodylus'' likely originated in Africa and radiated outwards towards Southeast Asia and the Americas, although an Australia/Asia origin has also been considered.
Phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
evidence supports ''Crocodylus'' diverging from its closest recent relative, the extinct ''
Voay ''Voay'' is an extinct genus of crocodile from Madagascar that lived during the Late Pleistocene to Holocene, containing only one species, ''V. robustus''. Numerous subfossils have been found, including complete skulls, noted for their distinctiv ...
'' of
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
, around 25 million years ago, near the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
/
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
boundary.


Phylogeny

Below is a
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
based on a 2018
tip dating Tip dating is a technique used in molecular dating that allows the inference of time-calibrated phylogenetic trees. Its defining feature is that it uses the ages of the samples to provide time information for the analysis, in contrast with traditio ...
study by Lee & Yates simultaneously using morphological, molecular (
DNA sequencing DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
), and
stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostra ...
(
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
age) data, as revised by the 2021 Hekkala ''et al.''
paleogenomics Paleogenomics is a field of science based on the reconstruction and analysis of genomic information in extinct species. Improved methods for the extraction of ancient DNA (aDNA) from museum artifacts, ice cores, archeological or paleontological site ...
study using DNA extracted from the extinct ''
Voay ''Voay'' is an extinct genus of crocodile from Madagascar that lived during the Late Pleistocene to Holocene, containing only one species, ''V. robustus''. Numerous subfossils have been found, including complete skulls, noted for their distinctiv ...
''.


Characteristics

Morelet's crocodile has a very broad snout with 66 to 68 teeth when they are fully mature. They are dark grayish-brown in color with dark bands and spots on the body and the tail. This is similar to other crocodiles, like the American crocodile, but the Morelet is somewhat darker. Juvenile crocodiles are bright yellow with some dark bands. The crocodile's iris is silvery brown. They have four short legs, giving them a rather sprawling gait, and a long tail, which is used for swimming. The hind feet of the crocodiles are webbed. They have very explosive capabilities because of their strong muscles and are fast runners.


Size

Morelet's crocodile is small compared to most other crocodiles. The males can become larger than the females. The average adult Morelet's crocodile is about , with a typical length range of (the lower measurement representing the mean total length of a female at sexual maturity which is attained at roughly 7–8 years of age in the wild).Platt, S. G., Rainwater, T. R., Thorbjarnarson, J. B., Finger, A. G., Anderson, T. A., & McMurry, S. T. (2009). ''Size estimation, morphometrics, sex ratio, sexual size dimorphism, and biomass of Morelet’s crocodile in northern Belize''. Caribbean Journal of Science, 45(1), 80-93. Almost all crocodiles in excess of are males and at this advanced stage of maturity, the male goes through a significant change in skull osteological morphology as the skull appears to increase in broadness and robustness. Large adult males can attain a length of , anything in excess of this is considered exceptionally rare for this species however the species has a maximum reported length of , with two other outsized specimens reportedly measuring , respectively. One mature adult specimen measuring and weighing had a bite force of . The weight of a large wild male crocodile is estimated to average around , although mass is likely much more in outsized individuals. One large male with a presumed total length of around weighed about . Overall, this species is similar in appearance and morphology to the
Cuban Cuban may refer to: * Something of, from, or related to Cuba, a country in the Caribbean * Cubans, people from Cuba, or of Cuban descent ** Cuban exile, a person who left Cuba for political reasons, or a descendant thereof * Cuban citizen, a perso ...
and the larger
American crocodile The American crocodile (''Crocodylus acutus'') is a species of crocodilian found in the Neotropics. It is the most widespread of the four extant species of crocodiles from the Americas, with populations present from South Florida and the coasts ...
s.


Distribution and habitat

Morelet's crocodile can be found in freshwater habitats in
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and along the
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...
stretching through Belize, Guatemala, and to Mexico.Dever, J. A; Richard E. Strauss, Thomas R. Rainwater, Scott T. McMurry, and Llewellyn D. Densmore III. “Genetic Diversity, Population Subdivision, and Gene Flow in Morelet’s Crocodile (Crocodylus moreletii) from Belize, Central America” Copeia. 2002. 4: 1078-1091. The
Belizean pine forests Belizean may refer to: * Something of, or related to Belize, a Central American nation * Belizean people, people originating in Belize whether they live there or in the Belizean diaspora * Belizean population, see Demographics of Belize * Belizean ...
are an example of the type of ecoregion in which they occur. In their freshwater habitats, they prefer isolated areas that are secluded. This species of crocodile can mainly be found in freshwater swamps and marshes which are located inland, and in large rivers and lakes. Both of these habitats are forested to help add cover. The Morelet can also be found along the coast in
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
waters and the grassy savannas on the
Yucatán Peninsula The Yucatán Peninsula (, also , ; es, Península de Yucatán ) is a large peninsula in southeastern Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north ...
. These crocodiles become much more distributed during the rainy seasons when flooding occurs and it is easier for them to move elsewhere. Juvenile crocodiles live in very dense cover to protect them from other predators that might be in the area and will remain there until they become older and able to fend for themselves. Adult crocodiles are known to dig out
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
s during
dry season The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The te ...
s in their area. The range of this crocodile can overlap with the American crocodile, which can sometimes lead to them being confused with one another. Recently, the Morelet's crocodile has been introduced into the
Rio Grande The Rio Grande ( and ), known in Mexico as the Río Bravo del Norte or simply the Río Bravo, is one of the principal rivers (along with the Colorado River) in the southwestern United States and in northern Mexico. The length of the Rio G ...
(Rio Bravo in Mexico). Several newspaper outlets on the Mexican side of the border report of reptiles inhabiting the river appearing not to be the
American alligator The American alligator (''Alligator mississippiensis''), sometimes referred to colloquially as a gator or common alligator, is a large crocodilian reptile native to the Southeastern United States. It is one of the two extant species in the g ...
which is native to
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
, but the Morelet's crocodile which is native to
Tamaulipas Tamaulipas (), officially the Free and Sovereign State of Tamaulipas ( es, Estado Libre y Soberano de Tamaulipas), is a state in the northeast region of Mexico; one of the 31 states which, along with Mexico City, comprise the 32 Federal Entiti ...
from
San Fernando San Fernando may refer to: People *Ferdinand III of Castile (c. 1200–1252), called ''San Fernando'' (Spanish) or ''Saint Ferdinand'', King of Castile, León, and Galicia Places Argentina *San Fernando de la Buena Vista, city of Greater Buenos ...
southward. Crocodiles have been seen in the cities of Matamoros,
Reynosa Reynosa () is a border city in the northern part of the state of Tamaulipas, in Mexico. It is also the municipal seat of Reynosa Municipality. The city is located on the southern bank of the Rio Grande in the international Reynosa–McAllen Metr ...
and as far north as
Nuevo Laredo Nuevo Laredo () is a city in the Municipality of Nuevo Laredo in the Mexican state of Tamaulipas. The city lies on the banks of the Rio Grande, across from Laredo, United States. The 2010 census population of the city was 373,725. Nuevo Lar ...
. The sightings have prompted several municipal police departments to put up signs warning people about entering the river.


Biology and behavior


Hunting and diet

Like most crocodilians, Morelet's crocodiles are highly opportunistic and will prey on practically anything that they can overpower that comes in their territories.Platt, S. G., Rainwater, T. R., Finger, A. G., Thorbjarnarson, J. B., Anderson, T. A., & McMurry, S. T. (2006). ''Food habits, ontogenetic dietary partitioning and observations of foraging behaviour of Morelet's crocodile (Crocodylus moreletii) in northern Belize''. The Herpetological Journal, 16(3), 281-290. Juvenile crocodiles feed largely on
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
and
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s until they become bigger and more capable of bringing down larger prey. Adults largely prey on small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s, other
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s, and fish, as well as
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
s,
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s and other invertebrates. These small mammals can include domesticated animals like cats and dogs. Large specimens have been shown to be able to overtake large prey.
Dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Do ...
s and
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s have been taken by this species, including a record of a adult killing an English sheepdog which weighed at least . Adults have also been recorded eating even larger animals, including adult and nearly-grown
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
and an adult
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk. Tapirs inhabit jungle and forest regions of South and Central America, with one species inhabit ...
, although these have been cases of scavenging on caracasses, with the tapir having been killed by a
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
(it is possible that the jaguar was present when the crocodile fed). Crocodiles have been known to be cannibalistic towards smaller specimens. Although previously thought to be inoffensive, there have been recent reports that the species has attacked humans on multiple occasions and at least 12 documented human fatalities have occurred. Despite the relatively small size of the species, large adult Morelet's are aggressive and easily capable of overpowering an unaware human near the water. Due to partial consumption, recorded fatal attacks are likely predatory rather than defensive in nature.


Reproduction

Breeding usually takes place between April and June and the eggs are laid before the start of the rainy season. Morelet's crocodiles are unique among North American crocodiles in that they build mound nests only, and not mound and hole nests. These mound nests are about wide and high and can be found near the water or on floating vegetation. A female crocodile can lay between 20 and 45 eggs and nests have been found containing eggs from more than one female. The eggs are buried and the nests are guarded by females. The eggs usually hatch after 80 days of incubation and hatchlings are normally about long. After the eggs have hatched the female crocodile will carry her young to the water where they are protected by both parents and will later leave them to fend for themselves. Females are highly protective of their young and have reportedly been observed to aggressively displace intruders and humans if distress calls of the baby crocodiles are heard and even father crocodiles have been observed to spring to the defense of young crocodiles. In captivity, juvenile crocodiles are treated aggressively by adult crocodiles but never hatchlings.


Conservation

Morelet's crocodile has long been threatened by habitat destruction and illegal hunting. Both of these factors have significantly lowered their populations. It was hunted for its hide during the 1940s and 1950s because high-quality leather can be made from their skins.Platt, Steven; John Thorbjarnarson. “Population status and conservation of Morelet’s Crocodile, Crocodylus moreletii, in northern Belize” Biological Conservation. 2000. 96: 21-29.


Notes


References


External links


"Crocodile Sighting in Corpus Christi"

"Species Name" Morelets Crocodile; Outcome: Fatal"
CrocBITE Live Feed * Britton, Adam

Crocodilian Conservation Database. 2002. {{Taxonbar, from=Q723708 Crocodylidae Reptiles of Mexico Reptiles of Belize Reptiles of Guatemala Reptiles of the Caribbean Least concern biota of Mexico Least concern biota of North America Reptiles described in 1851 Taxa named by Auguste Duméril Taxa named by Gabriel Bibron Crocodilians of North America