Moorland Clouded Yellow
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''Colias palaeno'', known by the common names moorland clouded yellow, palaeno sulphur, and pale Arctic clouded yellow, is a butterfly in the family Pieridae.


Subspecies

Subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
include:Funet
at Markku Savela's ''Lepidoptera and Some Other Life Forms''
* ''Colias palaeno palaeno'' –
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
, Norway, Finland, and Estonia * ''Colias palaeno aias'' Fruhstorfer, 1903
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
the largest form of ''palaeno'', the marginal band of the forewing being very broad and the underside rich green *''Colias palaeno baffinensis'' Ebner & Ferris, 978/small> – northern North America *''Colias palaeno chippewa'' W.H. Edwards, 1870 – northern North America, may be a separate species ('' C. chippewa'') * ''Colias palaeno europome'' (Esper, 1778)Belgium, Germany, Slovakia, Romania, and Ukraine * ''Colias palaeno europomene'' Ochsenheimer, 1808 – high altitude in the Alps. This form is somewhat larger than the nominate, the male lemon-yellow above, being below deeper yellow, as is also the female. * ''Colias palaeno orientalis'' Staudinger, 1892Kamchatka very similar to ''europomene'', being dark greenish on the underside of the hindwing * ''Colias palaeno poktusani'' O. Bang-Haas, 1934North Korea * ''Colias palaeno sachalinensis'' Matsumura, 1919Sakhalin * ''Colias palaeno synonyma'' Bryk, 1923
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
, Denmark''Colias''
at Markku Savela's ''Lepidoptera and Some Other Life Forms''
''Colias aias'' is treated as a full species by some authors.


Distribution

''Colias palaeno'' is a Holarctic species, widespread through Asia, Europe and North America. It is present in central and northern Europe from eastern France to the Baltic States and northern
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
, Norway, Finland and to the eastern Europe, then in eastern Siberia, in the Chukchi Peninsula, in
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
and in northern areas of North America.


Habitat

This species inhabits various types of moorland, Russian Insects
/ref> forest meadows, open coniferous forests and open areas which may contain scrub. In the southern areas of its range it is a high alpine species reaching an elevation over above sea level, but it can be found on upland bogs at an elevation of about . Though the species is normally restricted to these habitats, occasionally it is found far from suitable breeding grounds.


Description

''Colias palaeno'' has a wingspan of in males, of in females. Upperside of male pale yellow with blackish brown distal margins, pale-centred dark middle spot to the forewing and light middle pot to the hindwing; fringes red. Underside of forewing pale yellow with white-centred dark middle spot, the costal and distal edges being red; hindwing yellow, strongly dusted with fuscous, the large middle spot being mother-of-pearl colour, and the fringes red. The female has a white ground colour above, the underside of the forewing being white proximally, yellow at apex, the hindwing being somewhat paler in the female than in the male. The larva is sea-green, velvety, bearing minute black dots; a lateral stripe bright yellow edged with black beneath, below the same the white black-edged spiracles; underside and abdominal legs dull green, thoracic legs yellowish, head green. The Pupa is greenish yellow, the back being strongly convex.Kimmo Silvone
Larvae of North-European Lepidoptera
/ref> (MHNT) Colias palaeno - Majdan Kasztelański Pologne - male dorsal.jpg, ''Colias palaeno'' ♂ (MHNT) Colias palaeno - Majdan Kasztelański Pologne - male ventral.jpg, ''Colias palaeno'' ♂ △


Biology

It is a univoltine species, flying from June to August. As most ''Colias''-species, ''Colias palaeno'' is an avid flyer. The larva is found on bog bilberry ('' Vaccinium uliginosum''), on '' Vaccinium myrtillus'' and on ''
Vaccinium caespitosum ''Vaccinium cespitosum'' (also, ''caespitosum''), known as the dwarf bilberry, dwarf blueberry, or dwarf huckleberry, is a species of flowering shrub in the genus ''Vaccinium'', which includes blueberries, huckleberries, and cranberries. Descrip ...
''.


Gallery

Colias palaeno poktussani male Syntype 01 dorsal side ZSM.jpg, ''Colias palaeno poktussani ''. Male, dorsal side Colias palaeno poktussani female Syntype 01 dorsal side ZSM.jpg, ''Colias palaeno poktussani ''Female, dorsal side Colias palaeno poktussani male Syntype 01 ventral side ZSM.jpg, ''Colias palaeno poktussani ''Male, ventral side Colias palaeno poktussani female Syntype 01 ventral side ZSM.jpg, ''Colias palaeno poktussani ''Female, ventral side


References


External links

* Paolo Mazzei, Daniel Morel, Raniero Panfil
and Butterflies of Europe and North Africa



Butterflies and Moths of North America

Lepiforum

Lepi Net
Butterflies Along Alaska's Hiking Trails
''Colias palaeno (Linnaeus 1761) (Papilio)''
ZooBank {{Taxonbar, from=Q851578 palaeno Butterflies described in 1761 Butterflies of Asia Butterflies of Europe Butterflies of North America Insects of the Arctic Taxa named by Carl Linnaeus