Monarda Punctata; White Petals
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''Monarda'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s in the mint
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
,
Lamiaceae The Lamiaceae ( ) or Labiatae are a family (biology), family of flowering plants commonly known as the mint, deadnettle, or sage family. Many of the plants are aromatic in all parts and include widely used culinary herbs like basil (herb), ba ...
.Harley, R. M., et al. 2004. "Labiatae". pp 167-275 In: Kubitzki, K. (editor) and J. W. Kadereit (volume editor). ''The Families and Genera of Vascular Plants'' volume VII. Springer-Verlag: Berlin; Heidelberg, Germany. The genus is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to North America. Common names include bergamot, bee balm, horsemint, and oswego tea, the first being inspired by the fragrance of the leaves, which is reminiscent of
bergamot orange ''Citrus bergamia'', the bergamot orange (pronounced ), is a fragrant citrus fruit the size of an orange, with a yellow or green colour similar to a lime, depending on ripeness. Genetic research into the ancestral origins of extant citrus cul ...
(''Citrus bergamia''). The genus was named for the Spanish botanist
Nicolás Monardes Nicolás Bautista Monardes (1493 – 10 October 1588) was a Spanish physician and botanist. Monardes published several books of varying importance. In ''Diálogo llamado pharmacodilosis'' (1536), he examines humanism and suggests studying se ...
, who wrote a book in 1574 describing plants of the
New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...
.


Description

''Monarda'' species include
annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year **Yearbook **Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), a ...
and
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...
s. They grow erect to heights of . The slender, serrated,
lanceolate The following terms are used to describe leaf plant morphology, morphology in the description and taxonomy (biology), taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade ...
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are oppositely arranged on the square
stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...
,
hairless Hairless, also known as H, is a well-characterized Drosophila gene. Since Hairless is a dominant loss of function mutation, many mutations to Hairless are embryonic lethal, but there are several viable hairless mutants. This specific Drosophila ...
or sparsely hairy, and about long. The flowers are tubular and bilaterally symmetric, with a narrow upper lip and a wider lower lip. The wild flowers are single, but some cultivated forms have double flowers. They are
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...
, with male and female structures in each flower. There are two
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s.
Inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s occur at the top of the stem or emerge from the axils. They are typically crowded head-like clusters of flowers with leafy
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
s. Flower color varies, with wild species bearing red, pink, and light purple flowers. '' M. didyma'' has bright
carmine Carmine ()also called cochineal (when it is extracted from the Cochineal, cochineal insect), cochineal extract, crimson Lake pigment, lake, or carmine lake is a pigment of a bright-red color obtained from the aluminium coordination complex, compl ...
red flowers, '' M. fistulosa'' has pink, and '' M. citriodora'' and '' M. pectinata'' have pale purple. Hybrids occur in the wild, and they are common in cultivation. Seed collected from hybrids does not yield plants identical to the parent.


Uses

The crushed leaves of all species exude a spicy, fragrant
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the ...
. Of the species examined in one
study Study or studies may refer to: General * Education **Higher education * Clinical trial * Experiment * Field of study * Observational study * Scientific study * Research * Study skills, abilities and approaches applied to learning Other * Study ...
, ''M. didyma'' contained the highest concentration of oil. Several species, including ''
Monarda fistulosa ''Monarda fistulosa'', the wild bergamot or bee balm,Wild Bergamot
, Edmonton Naturalization Group ...
'' and '' M. didyma'', have a long history of use as medicinal plants by many Native Americans, such as the
Blackfoot The Blackfoot Confederacy, ''Niitsitapi'', or ''Siksikaitsitapi'' (ᖹᐟᒧᐧᒣᑯ, meaning "the people" or " Blackfoot-speaking real people"), is a historic collective name for linguistically related groups that make up the Blackfoot or Bl ...
,
Menominee The Menominee ( ; meaning ''"Menominee People"'', also spelled Menomini, derived from the Ojibwe language word for "Wild Rice People"; known as ''Mamaceqtaw'', "the people", in the Menominee language) are a federally recognized tribe of Na ...
,
Ojibwa The Ojibwe (; syll.: ᐅᒋᐺ; plural: ''Ojibweg'' ᐅᒋᐺᒃ) are an Anishinaabe people whose homeland (''Ojibwewaki'' ᐅᒋᐺᐘᑭ) covers much of the Great Lakes region and the northern plains, extending into the subarctic and thro ...
and Winnebago. The Blackfoot recognized the strong
antiseptic An antiseptic ( and ) is an antimicrobial substance or compound that is applied to living tissue to reduce the possibility of sepsis, infection, or putrefaction. Antiseptics are generally distinguished from ''antibiotics'' by the latter's abil ...
action of the plants, and used them in
poultice A poultice or cataplasm, also called a fomentation, is a soft moist mass, often heated and medicated, that is applied to the skin to reduce inflammation, soothe pain, promote healing, or otherwise treat wounds or ailments. Soft materials like cer ...
s for skin infections and minor wounds. Native Americans and later settlers also used it to alleviate stomach and bronchial ailments. A
tisane Herbal teas, technically known as herbal infusions, and less commonly called tisanes (UK and US , US also ), are beverages made from the infusion or decoction of herbs, spices, or other plant material in hot water. Often herb tea, or the plain ...
made from the plant was also used to treat mouth and throat infections caused by
dental caries Tooth decay, also known as caries,The word 'caries' is a mass noun, and is not a plural of 'carie'.'' is the breakdown of teeth due to acids produced by bacteria. The resulting cavities may be a number of different colors, from yellow to black ...
and
gingivitis Gingivitis is a non-destructive disease that causes inflammation of the gums; ulitis is an alternative term. The most common form of gingivitis, and the most common form of periodontal disease overall, is in response to bacterial biofilms (also ...
. Bee balm is a natural source of the antiseptic compound
thymol Thymol (also known as 2-isopropyl-5-methylphenol, IPMP), , is a toxic monoterpenoid phenol derivative of ''p''-Cymene, isomeric with carvacrol. It occurs naturally in the oil of thyme, and it is extracted from ''Thymus vulgaris'' (common thy ...
, the primary active ingredient in some modern commercial
mouthwash Mouthwash, mouth rinse, oral rinse, or mouth bath is a liquid which is held in the mouth passively or swirled around the mouth by contraction of the perioral muscles and/or movement of the head, and may be gargled, where the head is tilted back ...
formulas. The Winnebago used a bee balm tisane as a general
stimulant Stimulants (also known as central nervous system stimulants, or psychostimulants, or colloquially as uppers) are a class of drugs that increase alertness. They are used for various purposes, such as enhancing attention, motivation, cognition, ...
. Bee balm was also used as a
carminative A carminative, known in Latin as carminativum (plural carminativa), is a herb or preparation intended to combat flatulence either by preventing formation of gas in the gastrointestinal tract or Name The word ''carminative'' is a derivative of Lat ...
herb by Native Americans to prevent excessive
flatulence Flatulence is the expulsion of gas from the Gastrointestinal tract, intestines via the anus, commonly referred to as farting. "Flatus" is the medical word for gas generated in the stomach or bowels. A proportion of intestinal gas may be swal ...
.Tilford, G. L. ''Edible and Medicinal Plants of the West''. Although somewhat bitter due to the thymol content in the leaves and buds, the plant tastes like a mix of
spearmint Spearmint (''Mentha spicata''), also known as garden mint, common mint, lamb mint and mackerel mint, is native to Europe and southern temperate Asia, extending from Ireland in the west to southern China in the east. It is naturalized in many othe ...
and
peppermint Peppermint (''Mentha'' × ''piperita'') is a Hybrid (biology), hybrid species of Mentha, mint, a cross between Mentha aquatica, watermint and spearmint. Indigenous to Europe and the Middle East, the plant is now widely spread and cultivated in m ...
with
oregano Oregano (, ; ''Origanum vulgare'') is a species of flowering plant in the mint family, Lamiaceae. It was native to the Mediterranean region, but widely naturalised elsewhere in the temperate climate, temperate Northern Hemisphere. Oregano is a ...
. Bee balm was traditionally used by Native Americans as a
seasoning Seasoning is the process of supplementing food via herbs, spices, and/or salts, intended to enhance a particular flavour. General meaning Seasonings include herbs and spices, which are themselves frequently referred to as "seasonings". Salt may ...
for wild
game A game is a structured type of play usually undertaken for entertainment or fun, and sometimes used as an educational tool. Many games are also considered to be work (such as professional players of spectator sports or video games) or art ...
, particularly birds. The plants are widespread across North America and can be found in moist meadows, hillsides, and forest clearings up to in elevation.


Cultivation

''Monarda'' plants thrive in sun and moist but well-drained soil. Plants growing in partial shade spread horizontally and produce fewer flowers. ''Monarda'' are used in beds and borders to attract
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
s, pollinating insects, and insects that control garden pests. They are prone to developing
powdery mildew Powdery mildew is a fungus, fungal disease that affects a wide range of plants. Powdery mildew diseases are caused by many different species of Ascomycota, ascomycete fungi in the order Erysiphales. Powdery mildew is one of the easier plant disea ...
in high humidity, especially if planted in a place without good air circulation. Hybrid
cultivars A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated. Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting, tissue cult ...
range from dark red mahogany to bluish lilac to multiple shades of pink. These are generally not as robust as wild species. Some hybrids have been developed to produce high levels of essential oil for use as flavoring or medicine.


Cultivars

The following cultivars have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
: * 'Beauty of Cobham' (pink) * 'Gardenview Scarlet' * 'Marshall's Delight' (pink) * 'Squaw' (red) * 'Talud' (pink) The UK National Collection of Monardas is held at Glyn Bach Gardens at Pont Hywel, Efailwen, near
Llandissilio Llandissilio () is a village and parish in the community of Llandissilio West in east Pembrokeshire, Wales on the A478 road between Efailwen to the north and Clunderwen to the south. A largely ribbon development along the main road, the vill ...
in Pembrokeshire. Carole Whittaker of Glyn Bach Gardens was appointed
International Cultivar Registration Authority An International Cultivation Registration Authority (ICRA) is an organization responsible for ensuring that the names of plant cultivars and cultivar groups are defined and not duplicated. The ICRA system was established more 50 years ago, and op ...
for ''Mondarda'' in 2019, and currently recognises over 100 cultivars.


Ecology

''Monarda'' species are used as food plants by the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of some
Lepidoptera Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organ ...
species, including case-bearers of the genus ''
Coleophora ''Coleophora'' is a very large genus of moths of the family Coleophoridae. It contains some 1,350 described species. The genus is represented on all continents, but the majority are found in the Nearctic and Palaearctic regions. Many authors have t ...
''. '' Coleophora monardae'' feeds only on ''Monarda'' plants, and '' C. heinrichella'' and '' C. monardella'' only feed on the species ''M. fistulosa''.


Taxonomy

''Monarda'' is in the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Mentheae Mentheae is the largest tribe of plants in the family Lamiaceae. It includes herbs such as sage, hyssop, mint, bee balm and thyme. Genera Subtribe Lycopinae * '' Lycopus'' (21 living species) Subtribe Menthinae * '' Acanthomintha'' * '' Blephil ...
of the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Nepetoideae Nepetoideae is a subfamily of plants in the family Lamiaceae. , the Angiosperm Phylogeny Website (APweb) accepted the following genera: The format of the authorities is not the IPNI standard. * '' Acanthomintha'' (A. Gray) Bentham & J. D. Hooke ...
in the mint family.
Molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
studies of this tribe have been poorly sampled, and relationships within it remain unclear.Ryding, P. O. 2010. Pericarp structure and phylogeny of tribe Mentheae (Lamiaceae). ''Plant Systematics and Evolution'' 285(3-4), 165–75. The genera ''
Blephilia ''Blephilia'', the pagoda plant or wood mint, is a genus of four species of flowering plants in the family Lamiaceae. They are all herbaceous plants native to eastern North America. ''Blephilia'' are most often found in open areas, glades, and me ...
'' and ''
Pycnanthemum ''Pycnanthemum'' is a genus of herbaceous plants in the mint family (Lamiaceae). Species in this genus are often referred to as "mountain mints" and they often have a minty or thyme-like aroma when crushed. All species of ''Pycnanthemum'' are ...
'' are close relatives of ''Monarda'', but they might not be the closest. ''Monarda'' is divided into two distinct
subgenera In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
, ''Monarda'' and ''Cheilyctis''.Scora, R. W. 1967. Interspecific relationships in the genus ''Monarda'' (Labiatae). ''University of California Publications in Botany'' 41(1), 1–71. These are easily distinguished by several characteristics.


Species

Species in the genus include: * '' Monarda bartlettii'' Standl. - Tamaulipas, Veracuz * '' Monarda balmettii'' Nutt. - fools balm - northwest United States * '' Monarda bradburiana'' L.C.Beck – eastern beebalm - mid Mississippi Valley * ''
Monarda citriodora ''Monarda citriodora'' is a species of flowering plant in the mint family, Lamiaceae, that is native to the southern United States and northern Mexico. Common names include lemon beebalm, lemon mint (this may also apply to Eau de Cologne mint or ...
'' Cerv. ex Lag. – lemon beebalm, lemon-mint - southern United States, northern Mexico * '' Monarda clinopodia'' L. – white bergamot, basil beebalm - eastern United States, especially Appalachians * '' Monarda clinopodioides'' A.Gray – basil beebalm - Kansas, Oklahoma, Texas, Louisiana * ''
Monarda didyma ''Monarda didyma'', the crimson beebalm, scarlet beebalm, scarlet monarda, Eau-de-Cologne plant, Oswego tea, or bergamot, is a North American aromatic herb in the family Lamiaceae. Description ''M. didyma'' is a perennial plant that grows to i ...
'' L. – Oswego tea, scarlet beebalm, fragrantbalm, mountain-mint - eastern United States, especially Appalachians, eastern Canada * '' Monarda eplingiana'' Standl. - Coahuila * ''
Monarda fistulosa ''Monarda fistulosa'', the wild bergamot or bee balm,Wild Bergamot
, Edmonton Naturalization Group ...
'' L. – wild bergamot, mintleaf beebalm, horse-mint, purple beebalm - widespread across most of United States + Canada; Tamaulipas, Nuevo León; cultivated in China and elsewhereFlora of China Vol. 17 Page 223 拟美国薄荷 ni mei guo bo he ''Monarda fistulosa'' Linnaeus, Sp. Pl. 1: 22. 1753.
/ref> * '' Monarda fruticulosa'' Epling – spotted beebalm - southern Texas * '' Monarda humilis'' (Torr.) Prather & J.A.Keith - New Mexico * '' Monarda lindheimeri'' Engelm. & A.Gray ex A.Gray – Lindheimer's beebalm - Texas, Louisiana, southwestern Arkansas * '' Monarda luteola'' Singhurst & W.C.Holmes - northeastern Texas, southwestern Arkansas * '' Monarda maritima'' (Cory) Correll – seaside beebalm - coastal plain of Texas * ''
Monarda media ''Monarda media'' is a species of flowering plant in the mint family known by the common name purple bergamot. It is native to eastern North America, including the eastern United States and Ontario in Canada.Monarda × medioides'' W.H.Duncan - Georgia, Indiana ''(M. fistulosa × M. media)'' * ''
Monarda pectinata ''Monarda'' is a genus of flowering plants in the mint family, Lamiaceae.Harley, R. M., et al. 2004. "Labiatae". pp 167-275 In: Kubitzki, K. (editor) and J. W. Kadereit (volume editor). ''The Families and Genera of Vascular Plants'' volume VII. ...
'' Nutt. – plains beebalm, pony beebalm, spotted beebalm - central + southwestern United States (Great Plains, Rocky Mountains, southwestern desert mountains) * '' Monarda pringlei'' Fernald - Nuevo León * '' Monarda punctata'' L. – spotted beebalm, dotted monarda, horse-mint - Quebec, Ontario, eastern + south-central United States, California, northeastern Mexico * '' Monarda russeliana'' Nutt. ex Sims – redpurple beebalm - Texas, Oklahoma, Arkansas, Alabama, Mississippi, Kentucky * '' Monarda stanfieldii'' Small – Stanfield's beebalm - central Texas * '' Monarda viridissima'' Correll – green beebalm - east-central Texas


Formerly placed here

* ''
Blephilia ciliata ''Blephilia ciliata'' is a species of herbaceous perennial plant in the Lamiaceae (mint) family native to central and eastern North America. It is commonly called downy wood mint. Other common names include downy pagoda-plant, sunny woodmint and ...
'' (L.) Benth. (as ''M. ciliata'' L.) * ''
Blephilia hirsuta ''Blephilia hirsuta'', commonly known as hairy wood-mint or hairy pagoda plant, is a species of herbaceous perennial in the mint family Lamiaceae. It is native to eastern North America. Description Hairy wood mint is a perennial plant that i ...
'' (Pursh) Benth. (as ''M. ciliata'' Pursh)


References

; Secondary sources *


External links

* * {{Taxonbar, from=Q157665 Lamiaceae genera Taxa named by Carl Linnaeus Plants used in traditional Native American medicine Plants used in Native American cuisine