The molly miller (''Scartella cristata'') is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

of the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Blenniidae (

combtooth blennies Combtooth blennies are blenniiformids; percomorph marine fish of the family Blenniidae, part of the order Blenniiformes. They are the largest family of blennies with around 401 known species in 58 genera. Combtooth blennies are found in tropica ...

) in the

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

Perciformes. It can be found in the Atlantic, Mediterranean, and northwest Pacific Oceans. Its color is a

mottled Mottle is a pattern of irregular marks, spots, streaks, blotches or patches of different shades or colours. It is commonly used to describe the surface of plants or the skin of animals. In plants, mottling usually consists of yellowish spots o ...

tan, white, and black covering the body and fins. The head of this fish is covered with short hair-like appendages and has two very large eyes. This species reaches in

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

Reproduction

This blenny is

oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...

; its eggs are

demersal The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer of ...

and adhesive.

Habitat

The molly miller is a

marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...

tropical fish that lives in rocks or

coral reefs A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...

0–10 m below the surface. Coral reefs are perfect places for the fish to hunt and feed on the small

crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

and

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

that make up its omnivorous diet, and give them shelter and places to hide, as well.

Phylogeny

A 2020 study which analysed the

mitochondrial A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is use ...

data of the genus ''

Scartella ''Scartella'' is a genus of combtooth blennies found in the Atlantic and Indian oceans. Species There are currently seven recognized species in this genus: * '' Scartella caboverdiana'' Bath, 1990 * ''Scartella cristata'' ( Linnaeus, 1758) – M ...

'' for the first time showed that ''Scartella cristata'' is a lineage consisting of 5 clades: 2 in Caribbean waters, 1 in the East Atlantic/Mediterranean, and 2 in Brazil. The Brazilian clades are sympatric from

Rio de Janeiro Rio de Janeiro ( , , ; literally 'River of January'), or simply Rio, is the capital of the state of the same name, Brazil's third-most populous state, and the second-most populous city in Brazil, after São Paulo. Listed by the GaWC as a b ...

Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...

states (southern Brazil), with one clade being unique to Brazil and the other closely related to the eastern Atlantic lineage.

Distribution

It lives in the western

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...

from

Bermuda ) , anthem = "God Save the King" , song_type = National song , song = " Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , e ...

Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...

, and northern

Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

, and in the eastern Atlantic from

Mauritania Mauritania (; ar, موريتانيا, ', french: Mauritanie; Berber: ''Agawej'' or ''Cengit''; Pulaar: ''Moritani''; Wolof: ''Gànnaar''; Soninke:), officially the Islamic Republic of Mauritania ( ar, الجمهورية الإسلامية ...

and the

Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...

Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...

. The fish can be found in the southern

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...

, including near

Milan Milan ( , , Lombard: ; it, Milano ) is a city in northern Italy, capital of Lombardy, and the second-most populous city proper in Italy after Rome. The city proper has a population of about 1.4 million, while its metropolitan city h ...

Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...

, and the

Peloponnese The Peloponnese (), Peloponnesus (; el, Πελοπόννησος, Pelopónnēsos,(), or Morea is a peninsula and geographic regions of Greece, geographic region in southern Greece. It is connected to the central part of the country by the Isthmu ...

(

Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...

). It also lives near

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

and

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...

References

Bibliography

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San Francisco San Francisco (; Spanish language, Spanish for "Francis of Assisi, Saint Francis"), officially the City and County of San Francisco, is the commercial, financial, and cultural center of Northern California. The city proper is the List of Ca ...

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

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Massachusetts Massachusetts (Massachusett language, Massachusett: ''Muhsachuweesut assachusett writing systems, məhswatʃəwiːsət'' English: , ), officially the Commonwealth of Massachusetts, is the most populous U.S. state, state in the New England ...

. * Moyle, P. i J. Cech. (2000). ''Fishes: An Introduction to Ichthyology'', 4a. edició, Upper Saddle River,

Nova Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware Ri ...

: Prentice-Hall. * Nelson, J.S., E.J. Crossman, H. Espinosa-Pérez, L.T. Findley, C.R. Gilbert, R.N. Lea i J.D. Williams, 2004. Common and scientific names of fishes from the United States, Canada, and Mexico. American Fisheries Society, Special Publication 29, Bethesda,

Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...

. * Nelson, Joseph S.: ''Fishes of the World'', John Wiley & Sons. . 2006. * Nieder, J. (1988). Zum Vorkommen von ''Scartella cristata'' (L.) und ''Parablennius pilicornis'' (Cuv.) (Teleostei, Blenniidae) an der nordspanischen Mittelmeerküste. ''Zool. Anz.'' 220(3/4):144-150. * Nieder, J. (1996). Agonistic behavioural patterns in the blenniid fish ''Scartella cristata'' (Teleostei, Blenniidae) and the role of its unusual erectile nuchal crest. ''Z. Fischkd.'' 3(2):235-241. * Nieder, J. (1997). Seasonal variation in feeding patterns and food niche overlap in the Mediterranean blennies ''Scartella cristata'', ''Parablennius pilicornis'' and ''Lipophrys trigloides'' (Pisces: Blenniidae). ''Mar. Ecol.'' 18(3):227-237. * Nieder, J., G. La Mesa & M. Vacchi (2000). Blennidae along the Italian coasts of the Ligurian and the Tyrrhenian Sea: community structure and new records of ''Scartella cristata'' for northern Italy. ''Cybium'' 24(4):359-369. * Osório, B. (1898). Da distribuição geográfica dos peixes e crustáceos colhidos nas possessões portuguesas da África Occidental e existentes no Museu Nacional de Lisbon. ''J. Sci. Math. Phys. Nat.'' 5(19):185-207. * Rangel, C. A. & L. F. Mendes (2009). Review of blenniid fishes from Fernando de Noronha Archipelago, Brazil, with description of a new species of ''Scartella'' (Teleostei: Blenniidae). ''Zootaxa Núm.'' 2006: 51–61. * Springer, V. G. (1986). Blenniidae. p. 742–755. A. M. M. Smith & P. C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag,

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San Diego San Diego ( , ; ) is a city on the Pacific Ocean coast of Southern California located immediately adjacent to the Mexico–United States border. With a 2020 population of 1,386,932, it is the List of United States cities by population, eigh ...

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{{Taxonbar, from=Q1899882 Molly Miller Fish described in 1758 Taxa named by Carl Linnaeus Fish of the Atlantic Ocean