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Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with
broadleaf tree A broad-leaved, broad-leaf, or broadleaf tree is any tree within the diverse botanical group of angiosperms that has flat leaves and produces seeds inside of fruits. It is one of two general types of trees, the other being a conifer, a tree with n ...
ecoregions, and with
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
and broadleaf tree mixed coniferous forest ecoregions. These forests are richest and most distinctive in central China and eastern
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, with some other globally distinctive ecoregions in the Caucasus, the Himalayas, Southern Europe, Australasia, Southwestern South America and the Russian Far East.


Ecology

The typical structure of these forests includes four layers. * The uppermost layer is the canopy composed of tall mature trees ranging from high. Below the canopy is the three-layered,
shade-tolerant In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial ...
understory that is roughly shorter than the canopy. * The top layer of the understory is the sub-canopy composed of smaller mature trees, saplings, and suppressed juvenile canopy layer trees awaiting an opening in the canopy. * Below the sub-canopy is the
shrub layer Stratification in the field of ecology refers to the vertical layering of a habitat; the arrangement of vegetation in layers. It classifies the layers (sing. ''stratum'', pl. ''strata'') of vegetation largely according to the different heights to w ...
, composed of low growing woody plants. * Typically the lowest growing (and most
diverse Diversity, diversify, or diverse may refer to: Business *Diversity (business), the inclusion of people of different identities (ethnicity, gender, age) in the workforce * Diversity marketing, marketing communication targeting diverse customers ...
) layer is the ground cover or
herbaceous layer Stratification in the field of ecology refers to the vertical layering of a habitat; the arrangement of vegetation in layers. It classifies the layers (sing. ''stratum'', pl. ''strata'') of vegetation largely according to the different heights to w ...
.


Trees

In the Northern hemisphere, characteristic dominant
broadleaf tree A broad-leaved, broad-leaf, or broadleaf tree is any tree within the diverse botanical group of angiosperms that has flat leaves and produces seeds inside of fruits. It is one of two general types of trees, the other being a conifer, a tree with n ...
s in this biome include oaks ('' Quercus'' spp.), beeches ('' Fagus'' spp.), maples ('' Acer'' spp.), or birches ('' Betula'' spp.). The term "mixed forest" comes from the inclusion of coniferous trees as a canopy component of some of these forests. Typical coniferous trees include pines ('' Pinus'' spp.), firs ('' Abies'' spp.), and spruces (''
Picea A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
'' spp.). In some areas of this biome, the
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
s may be a more important canopy species than the broadleaf species. In the Southern Hemisphere, endemic genera such as '' Nothofagus'' and '' Eucalyptus'' occupy this biome, and most coniferous trees (members of the Araucariaceae and Podocarpaceae) occur in mixtures with broadleaf species, and are classed as broadleaf and mixed forests.


Climate

Temperate broadleaf and mixed forests occur in areas with distinct warm and cool seasons that give them moderate annual average temperatures — . These forests occur in relatively warm and rainy climates, sometimes also with a distinct dry season. A dry season occurs in the winter in East Asia and in summer on the wet fringe of the Mediterranean climate zones. Other areas, such as central eastern North America, have a fairly even distribution of rainfall; annual rainfall is typically over and often over , though it can go as low as in some parts of the Middle East and close to in the mountains of New Zealand and the Azores. Temperatures are typically moderate except in parts of Asia such as
Ussuriland Primorsky Krai (russian: Приморский край, r=Primorsky kray, p=prʲɪˈmorskʲɪj kraj), informally known as Primorye (, ), is a federal subject (a krai) of Russia, located in the Far East region of the country and is a part of the ...
, where temperate forests can occur despite very harsh conditions with very cold winters. The climates are typically humid for much of the year, usually appearing in the
humid subtropical climate A humid subtropical climate is a zone of climate characterized by hot and humid summers, and cool to mild winters. These climates normally lie on the southeast side of all continents (except Antarctica), generally between latitudes 25° and 40° ...
without summer being extremely hot and winter very mild, and in the humid continental climate zones to the south of tundra and the generally
subarctic The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Generally, ...
taiga. In the Köppen climate classification they are represented respectively by '' Cfa'', '' Dfa/Dfb'' southern range and '' Cfb'', and more rarely, '' Csb'', '' BSk'' and ''
Csa CSA may refer to: Arts and media * Canadian Screen Awards, annual awards given by the Academy of Canadian Cinema & Television * Commission on Superhuman Activities, a fictional American government agency in Marvel Comics * Crime Syndicate of Amer ...
''.


Ecoregions


Australasia


Eurasia


Americas


See also

* Mixed coniferous forest * Kuchler plant association system *
Mediterranean forests, woodlands, and scrub Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in ...
* Temperate deciduous forest *
Trees of the world A listing of lists of trees. * List of individual trees, including actual and mythical trees * List of largest giant sequoias * List of old growth forests * List of oldest trees * List of superlative trees **List of superlative trees in Sweden * ...


References


External links


World Wildlife Fund−WWF Biomes: Temperate broadleaf and mixed forests biome


* ttps://web.archive.org/web/20070310230140/http://forest.terraformers.ca/ Terraformers Canadian Forest Conservation Foundation {{Biomes Forests Terrestrial biomes