Missulena
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''Missulena'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
mygalomorph The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to t ...
spiders in the family
Actinopodidae Actinopodidae ( Mouse spiders) is a family of mygalomorph orb weaver spiders found in mainland Australia and South America usually in open forest. Species are most common in Queensland, Australia. It includes mouse spiders, whose bites, though r ...
. It was first described by Charles Walckenaer in 1805, and is a
senior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnae ...
of ''Eriodon''. ''M. tussulena'' is found in
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, but the rest are indigenous to Australia. They are sometimes referred to as "mouse spiders" from the now-disproven belief that they dig deep burrows similar to those of mice. ''
Scotophaeus blackwalli ''Scotophaeus blackwalli'', also known as the mouse spider, is a species of spider belonging to the family Gnaphosidae. Description The adult males of these spiders reach 9mm in length, maturing in the early summer, while females reach 12mm, and ...
'' is also called a "mouse spider", but it is smaller and not closely related.


Description

These spiders are medium to large in size, ranging from . They have a glossy carapace and high, broad heads with eyes spread out across the front of the head, and short
spinnerets A spinneret is a silk-spinning organ of a spider or the larva of an insect. Some adult insects also have spinnerets, such as those borne on the forelegs of Embioptera. Spinnerets are usually on the underside of a spider's opisthosoma, and are ...
in the rear of the
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the to ...
. They mostly
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...
s, though they may consume other small animals as opportunity presents. Their primary predators include
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. ...
s, centipedes, and
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always en ...
s. These spiders also exhibit
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
. Females are entirely black, while males coloration is specific to each species. For instance, male eastern mouse spiders (''M. bradleyi'') have a bluish patch, while male red-headed mouse spiders (''M. occatoria'') are brownish or blue-black with bright red-tinged jaws.


Identification

They resemble most genera of the infraorder
Mygalomorphae The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to the ...
. But they can be easily distinguished by the large pair of
chelicerae The chelicerae () are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as " jaws", chelicerae may be shaped as either articulated fangs, or similarl ...
, their two small eyes in the center and 3 to the sides. While in all other trap door spiders they are grouped in a mound at the center of the head. Females are harder to identify than males, as females are entirely black, while males may be brighter in color. Usually in form of a blueish abdomen or reddish carapace and chelicerae, or both.


Habitat and range

These spiders have a Gondwanan distribution, with one species found in
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
and the rest distributed throughout Australia. They live in trapdoor covered burrows that can extend to nearly in depth. Females generally remain in their burrows, depending on the males to wander in search of mates.


Medical significance

Bites from these spiders are painful, but not generally dangerous. Serious
envenomation Envenomation is the process by which venom is injected by the bite or sting of a venomous animal. Many kinds of animals, including mammals (e.g., the northern short-tailed shrew, ''Blarina brevicauda''), reptiles (e.g., the king cobra), spiders ...
s are relatively rare, but bites that have been documented in medical literature didn't require antivenom treatment or involve serious symptoms. There is evidence to suggest that a mouse spider bite can potentially be as serious as that of an Australian funnel-web spider, but recorded bites are rare, despite the abundance of some species amid human habitation. These spiders look very similar to the Australian funnel web spiders and bites should be initially treated as funnel web spider bites until the spider is positively identified by an expert. Australian funnel-web spider antivenom has been found to be effective in treating severe mouse spider bites. Unlike the Australian funnel-web spiders, however, the mouse spider is far less aggressive towards humans, and may often bite without releasing any venom.


Species

it contains 21 species: *'' M. bradleyi'' Rainbow, 1914 – Australia (New South Wales) *'' M. davidi'' Greenberg, Huey, Framenau & Harms, 2021 – Australia (Western Australia) *'' M. dipsaca'' Faulder, 1995 – Australia *'' M. faulderi'' Harms & Framenau, 2013 – Australia (Western Australia) *'' M. granulosa'' (O. Pickard-Cambridge, 1869) – Australia (Western Australia) *'' M. harewoodi'' Framenau & Harms, 2017 – Australia (Western Australia) *'' M. hoggi'' Womersley, 1943 – Australia (Western Australia) *'' M. insignis'' (O. Pickard-Cambridge, 1877) – Australia *'' M. iugum'' Greenberg, Huey, Framenau & Harms, 2021 – Australia (Western Australia) *'' M. langlandsi'' Harms & Framenau, 2013 – Australia (Western Australia) *'' M. leniae'' Miglio, Harms, Framenau & Harvey, 2014 – Australia (Western Australia) *'' M. mainae'' Miglio, Harms, Framenau & Harvey, 2014 – Australia (Western Australia) *'' M. manningensis'' Greenberg, Huey, Framenau & Harms, 2021 – Australia (Western Australia) *'' M. melissae'' Miglio, Harms, Framenau & Harvey, 2014 – Australia (Western Australia) *'' M. occatoria'' Walckenaer, 1805 – Southern Australia *'' M. pinguipes'' Miglio, Harms, Framenau & Harvey, 2014 – Australia (Western Australia) *'' M. pruinosa'' Levitt-Gregg, 1966 – Australia (Western Australia, Northern Territory) *'' M. reflexa'' Rainbow & Pulleine, 1918 – Australia (South Australia) *'' M. rutraspina'' Faulder, 1995 – Australia (Western Australia, South Australia, Victoria) *'' M. torbayensis'' Main, 1996 – Australia (Western Australia) *'' M. tussulena'' Goloboff, 1994 – Chile Formerly included: *''M. bonneti'' (Zapfe, 1961) (Transferred to '' Plesiolena'')


See also

* List of Actinopodidae species


References


Further reading

* * * * * * {{Taxonbar, from=Q100898 Mygalomorphae genera Taxa named by Charles Athanase Walckenaer Spiders of Australia Spiders of South America Fauna of Chile