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''Milnesium'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
tardigrade Tardigrades (), known colloquially as water bears or moss piglets, are a phylum of eight-legged segmented micro-animals. They were first described by the German zoologist Johann August Ephraim Goeze in 1773, who called them Kleiner Wasserbär ...
s. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
, the oldest species found so far (''M. swolenskyi'') is known from
Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...
stage deposits on the east coast of the United States.


Species

Described species include: * ''
Milnesium alabamae ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from ...
'' Wallendorf & Miller, 2009 *'' Milnesium almatyense'' Tumanov, 2006 * ''
Milnesium alpigenum ''Milnesium alpigenum'' is a species of tardigrade that falls under the Tardigrade, Tardigrada phylum. Like its Taxonomy, taxonomic relatives it is an Omnivore, omnivorous Predation, predator that feeds on other small organisms, such as algae, rot ...
'' Ehrenberg, 1853 *'' Milnesium antarcticum'' Tumanov, 2006 * ''
Milnesium argentinum ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Roszkowska, Ostrowska & Kaczmarek, 2015 *''
Milnesium asiaticum ''Milnesium asiaticum'' is a species of Eutardigrades in the family Milnesiidae. This species differs from its cogenerate species mainly by proportions of its claws and buccopharyngeal apparatus. References Further reading

* Apochela ...
'' Tumanov, 2006 * '' Milnesium barbadosense'' Meyer & Hinton, 2012 * ''
Milnesium beasleyi ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Kaczmarek, Jakubowska & Michalczyk, 2012 * ''
Milnesium beatae ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Roszkowska, Ostrowska & Kaczmarek, 2015 * '' Milnesium berladnicorum'' Ciobanu, Zawierucha, Moglan & Kaczmarek, 2014 * '' Milnesium bohleberi'' Bartels, Nelson, Kaczmarek & Michalczyk, 2014 *''
Milnesium brachyungue ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Binda & Pilato, 1990 * ''
Milnesium cassandrae ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Moreno-Talamantes, Roszkowska, García-Aranda, Flores-Maldonado & Kaczmarek, 2019 *''
Milnesium burgessi ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Schlabach, Donaldson, Hobelman, Miller & Lowman, 2018 * ''
Milnesium dornensis ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Ciobanu, Roszkowska & Kaczmarek, 2015 *''
Milnesium dujiangensis ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Yang, 2003 *''
Milnesium eurystomum ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Maucci, 1991 * ''
Milnesium fridae ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Moreno-Talamantes, León-Espinosa, García-Aranda, Flores-Maldonado & Kaczmarek, 2020 * ''
Milnesium granulatum ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Ramazzotti, 1962 * ''
Milnesium jacobi ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Meyer & Hinton, 2010 *''
Milnesium katarzynae ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Kaczmarek, Michalczyk & Beasley, 2004 * ''
Milnesium kogui ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Londoño, Daza, Caicedo, Quiroga & Kaczmarek, 2015 * ''
Milnesium krzysztofi ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Kaczmarek & Michalczyk, 2007 * '' Milnesium lagniappe'' Meyer, Hinton & Dupré, 2013 *'' Milnesium longiungue'' Tumanov, 2006 * '' Milnesium minutum'' Pilato & Lisi, 2016 * '' Milnesium quadrifidum'' Nederström, 1919 *'' Milnesium reductum'' Tumanov, 2006 *'' Milnesium reticulatum'' Pilato, Binda & Lisi, 2002 * '' Milnesium sandrae'' Pilato & Lisi, 2016 * ''
Milnesium shilohae ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Meyer, 2015 *''
Milnesium swansoni ''Milnesium'' is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (''M. swolenskyi'') is known from T ...
'' Young, Chappell, Miller & Lowman, 2016 *† '' Milnesium swolenskyi'' Bertolani & Grimaldi, 2000 *''
Milnesium tardigradum ''Milnesium tardigradum'' is a cosmopolitan species of tardigrade that can be found in a diverse range of environments. It has also been found in the sea around Antarctica. ''M. tardigradum'' was described by Louis Michel François Doyère in 184 ...
'' Doyère, 1840 *'' Milnesium tetralamellatum'' Pilato & Binda, 1991 * '' Milnesium tumanovi'' Pilato, Sabella & Lisi, 2016 * '' Milnesium validum'' Pilato, Sabella, D'Urso & Lisi, 2017 * '' Milnesium variefidum'' Morek, Gąsiorek, Stec, Blagden & Michalczyk, 2016 * '' Milnesium vorax'' Pilato, Sabella & Lisi, 2016 * '' Milnesium zsalakoae'' Meyer & Hinton, 2010


References

Apochela Tardigrade genera {{Tardigrade-stub