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Mildew is a form of
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from th ...
. It is distinguished from its closely related counterpart, mould, largely by its colour: moulds appear in shades of black, blue, red, and green, whereas mildew is white. It appears as a thin, superficial growth consisting of minute hyphae (
fungal A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from th ...
filaments) produced especially on living plants or organic matter such as wood, paper or leather. Both mould and mildew produce distinct offensive odours, and both have been identified as the cause of certain human ailments. In horticulture, mildew is either species of
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from th ...
in the order
Erysiphales Erysiphales are an order of ascomycete fungi. The order contains one family, Erysiphaceae. Many of them cause plant diseases called powdery mildew. Systematics The order contains one family (Erysiphaceae), 28 genera and approximately 100 speci ...
, or fungus-like organisms in the family ''
Peronosporaceae Peronosporaceae are a family of water moulds that contains 21 genera, comprising more than 600 species. Most of them are called downy mildews. Peronosporaceae are obligate biotrophic plant pathogens. They parasitise their host plants as an ...
''. It is also used more generally to mean
mould growth Mold (American English) or mould (British English), also sometimes referred to as mildew, is a fungal growth that develops on wet materials. Mold is a natural part of the environment and plays an important part in nature by breaking down dead or ...
. In
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
, mildew meant honeydew (a substance secreted by
aphid Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A t ...
s on leaves, formerly thought to distill from the air like
dew Dew is water in the form of droplets that appears on thin, exposed objects in the morning or evening due to condensation. As the exposed surface cools by radiating its heat, atmospheric moisture condenses at a rate greater than that at wh ...
), and later came to mean mould or fungus. Mildew grows on damp cloth, leather, or on plants, and growing on leaves can damage the plant.


Household varieties

The term mildew is often used generically to refer to
mould growth Mold (American English) or mould (British English), also sometimes referred to as mildew, is a fungal growth that develops on wet materials. Mold is a natural part of the environment and plays an important part in nature by breaking down dead or ...
, usually with a flat growth habit. Moulds can thrive on many organic materials, including clothing, leather, paper, and the ceilings, walls and floors of homes or offices with poor moisture control. Mildew can be cleaned using specialized mildew remover, or substances such as bleach (though they may discolour the surface). There are many species of mould. The black mould which grows in attics, on window sills, and other places where moisture levels are moderate often is ''
Cladosporium ''Cladosporium'' is a genus of fungi including some of the most common indoor and outdoor molds. Species produce olive-green to brown or black colonies, and have dark-pigmented conidia that are formed in simple or branching chains. Many species ...
''. Colour alone is not always a reliable indicator of the species of mould. Proper identification requires a microbiologist or mycologist. Mould growth found on cellulose-based substrates or materials where moisture levels are high (90 percent or greater) is often '' Stachybotrys chartarum.'' "Black mould," also known as "toxic black mould", properly refers to ''S. chartarum''. This species is commonly found indoors on wet materials containing cellulose, such as wallboard (drywall), jute, wicker, straw baskets, and other paper materials. ''S. chartarum'' does not, however, grow on plastic, vinyl, concrete, glass, ceramic tile, or metals. A variety of other mould species, such as ''
Penicillium ''Penicillium'' () is a genus of ascomycetous fungi that is part of the mycobiome of many species and is of major importance in the natural environment, in food spoilage, and in food and drug production. Some members of the genus produce pe ...
'' or ''
Aspergillus ' () is a genus consisting of several hundred mold species found in various climates worldwide. ''Aspergillus'' was first catalogued in 1729 by the Italian priest and biologist Pier Antonio Micheli. Viewing the fungi under a microscope, Mic ...
'', may appear to grow on non-cellulosic surfaces but are actually growing on the
biofilm A biofilm comprises any syntrophic consortium of microorganisms in which cells stick to each other and often also to a surface. These adherent cells become embedded within a slimy extracellular matrix that is composed of extracellular ...
that adheres to these surfaces. Glass, plastic, and concrete provide no food for organic growth and as such cannot support mould or mildew growth alone without biofilm present. In places with stagnant air, such as basements, moulds can produce a strong musty odour. The pink "mildew" often found on plastic shower curtains and bathroom tile is actually a red yeast, ''
Rhodotorula ''Rhodotorula'' is a genus of pigmented yeasts, part of the division Basidiomycota. It is readily identifiable by distinctive orange/red colonies when grown on Sabouraud's dextrose agar (SDA). This distinctive color is the result of pigme ...
''.


Environmental conditions

Mildew requires certain factors to develop. Without any one of these, it cannot reproduce and grow. The requirements are a food source (any organic material), sufficient ambient moisture (a
relative humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity dep ...
of between 62 and 93 percent), and reasonable warmth ( to is optimal, but some growth can occur anywhere between freezing and ). Slightly acidic conditions are also preferred. At warmer temperatures, air is able to hold a greater volume of water; as air temperatures drop, so does the ability of air to hold moisture, which then tends to condense on cool surfaces. This can work to bring moisture onto surfaces where mildew is then likely to grow (such as an exterior wall). Preventing the growth of mildew therefore requires a balance between moisture and temperature. This can be achieved by minimizing the moisture available in the air. Air temperatures at or below will inhibit growth, but only if the
relative humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity dep ...
is low enough to prevent water condensation (i.e., the
dew point The dew point is the temperature to which air must be cooled to become saturated with water vapor, assuming constant air pressure and water content. When cooled below the dew point, moisture capacity is reduced and airborne water vapor will cond ...
is not reached). With warmer temperatures, the water holding capacity of the air increases. This means that if the amount of water vapour in the warming air remains the same, the air will become drier (i.e. it has a lower
relative humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity dep ...
). This again inhibits fungal growth. Warm, growth-favoring temperatures coupled with ''high'' relative humidity, however, will set the stage for mildew growth. Air conditioners are one effective tool for removing moisture and heat from otherwise humid warm air. The coils of an air conditioner cause moisture in the air to condense on them, eventually losing this excess moisture through a drain and placing it back into the environment. They can also inhibit mildew growth by lowering indoor temperatures. In order for them to be effective, air conditioners must recirculate the existing indoor air and not be exposed to warm, humid outside air. Some energy efficient air conditioners may cool a room so quickly that they do not have an opportunity to also effectively collect and drain significant ambient water vapour.


See also

*
Downy mildew Downy mildew refers to any of several types of oomycete microbes that are obligate parasites of plants. Downy mildews exclusively belong to the Peronosporaceae family. In commercial agriculture, they are a particular problem for growers of cruc ...
* Phase I environmental site assessment *
Powdery mildew Powdery mildew is a fungal disease that affects a wide range of plants. Powdery mildew diseases are caused by many different species of ascomycete fungi in the order Erysiphales. Powdery mildew is one of the easier plant diseases to identify, a ...
* Obligate parasite


References

{{Reflist Building defects Plant pathogens and diseases